Craspedophorus tetrastigma tetrastigma ( Chaudoir, 1850 )

Häckel, Martin, 2017, A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae), Zootaxa 4330 (1), pp. 1-67 : 27-28

17a. Craspedophorus tetrastigma tetrastigma ( Chaudoir, 1850)

( Plate 3 , Figs 18–19, Plate 12 , Fig. 96)

Epicosmus tetrastigma Chaudoir, 1850: 417 (type locality “ Guinée et Sénégal [= Guinea, Senegal]”). Schaum 1853: 435, Chaudoir 1861: 339. Eudema tetrastigma Gemminger and Harold 1868: 210 . Craspedophorus tetrastigma Chaudoir 1879: 91 . Lorenz 2005: 321. Häckel and Farkač 2012: 83.

Isotarsus sinuaticollis Laferté-Sénectere 1851: 221 (type locality “Guin. Lusit. [=Guinea Bissau]”). Panagaeus sinuatocollis Chaudoir 1861: 339 [in error] syn nov.

Type material. Lectotype (♂), established by Chaudoir (1879: 91): “ SinuatÏcollis / Guin. Lusit. [handwritten in black on blue upper pinned label]// ♂ [handwritten in black on white second upper pinned label]// Ex Musaeo / Chaudoir [printed in red on white third upper pinned label]// Muséum Paris / ex Coll. / R. Oberthür / 1952 [printed in black on blue lower pinned label]// Type [printed in black on red lowest pinned label]”/// pinned first from left in first row of the series labelled: “ tetrastigma / Chaud. / Sénégal / 48. Buquet [handwritten in black on white box label in Chaudoir's Collection]” ( Plate 3 , Fig. 18, MNHN) . Paralectotypes. 1♂: “Ex Musaeo / Chaudoir [printed in red on white upper pinned label]// Muséum Paris / ex Coll. / R. Oberthür / 1952 [printed in black on blue lower pinned label]”; 1 (unknown sex): “Ex Musaeo / Chaudoir [printed in red on white upper pinned label]// Muséum Paris / ex Coll. / R. Oberthür / 1952 [printed in black on blue lower pinned label]”/// pinned second and third from left in first row of the same series in Chaudoir's Collection ( MNHN).

Additional material examined. Guinea. 1♂ labeled as compared type by Basilewsky: “ Beyla ” ( Plate 12 , Fig. 96, NMPC) . 1♂, 1♀ labeled as C. strachani (Hope) by Basilewsky: “Beyla” (cJB). Senegal. 1♀: “ Sénégambie // Ex Musaeo / Mniszech // Muséum Paris / ex Coll. / R. Oberthür / 1952” ( Plate 3 , Fig. 19, Chaudoir's Collection, MNHN) .

Note. The species was firstly based on several specimens in Buquet‘s collection. Description (in part, see Chaudoir 1850 /1: 417). “Length 10 points [= 20 mm]. This nice insect is distinctly different from Panagoeus regalis (Gory, 1833) ( grandis Buquet ). It differs in its somewhat smaller statue. Head less elongated, less protruded and not narrowing behind eyes; frontal grooves less deeply impressed, less grossly punctured; palps and antennae more slender, antennae less long, intermediate articles less compressed and dilated. Pronotum with anterior margin less sinuate; base almost flat; very weakly excavated near each lateral margin and posterior angle, which markedly more distinct, also not too obtuse and foregone by more distinct excavation towards angle; lateral rim elevated equally [as in C. regalis ], but herewith less convex in its posterior half; sagittal line more distinct; dorsum less grossly chagrined, more punctured; more finelly punctured laterally, on the convex part of lateral rim. Elytra strongly shorter, more ovoid; elytral margins more rounded as well as humeri, not dilated laterally; disk less convex, more covered by setae mainly anteriorly; striae less strongly punctured; intervals as well more weakly and sparsely punctured. Underside as well less punctured, mainly near prosternal margins and ventrally. Elytra as well ornate with four large maculae, but yellow colour of each macula is ligther, more as lemon, maculae less tranverse, humeral macula closer to base and elevated to humerus; preapical not narrowing laterally. I have compared two specimens of this species with two of Panagaeus regalis and consider these differences constant. It occurs in Guinea and Senegal. I am due to Mr. Buquet” [from French]. LaFerté-Sénectere added a brief key to species, for his new species Isotarsus sinuaticollis he noted (1851: 221). “(Prémiére division): larger species. Antennae longer than midlength of the body, distal attenuated, 3rd article twice than the second. Legs black; (aa) pronotum not convex, margins moreless elevated, (bb) transverse, not too annular, with sinuate margins, (cc) without strangulation at base (peduncle)” [translated from the French original]. Chaudoir (1861: 339) announces that he acquired three specimens from Bocandé's collection (i. e. described by LaFerté-Sénectere, 1851 as Isotarsus sinuaticollis coming from Portuguese territories on the northern coast of Guinea Bay, labeled by Laferté „Gui. Lusit.“). Chaudoir synonymizes Laferté‘s taxon with his species Epicosmus tetrastigma , the name [ sinuatocollis ] is wrongly typed in the text. Chaudoir later expands Laferté‘s brief description and establishes his lectotypes from this series. ( Chaudoir 1879: 91). “Length 20–21 mm, width 8.5 mm. Head about one third longer than ist frontal width before eyes, which strongly convex; frons chagrined and punctured, weakly depressed beside each margin; epistoma smooth, weakly convex, with impressions very arcuate towards base, near each margin; frons and vertex with transversal depression, distinctly visible, rugate, rugosity cumulated anteriorly on vertex, vertical base smooth. Pronotum twice or less wider than head, less long than wide, but not too transverse, hexagonal-shaped, with equal width of both sides, less enlarged at midlength, maximal width not at midlength, anterior margin less sinuate, anterior angles not removed from neck, weakly protruded, but widely rounded; lateral margins strongly rounded, mainly at midlength, from where sinuate posteriorly, longly, quite distinctly; with a small indentation in front of posterior angles, each with a dent, small, obtuse; base sinuate near each peduncular margin, and turned posteriorly to hind angles; dorsum quite grossly, not densely punctured; disk weakly convex, separated from lateral rims by a wide gutter, which more farther from margins posteriorly; lateral rims elevated, mainly posteriorly, but visible also along anterior half of lateral margin, although very fine here, and however long, very weakly marked; less grossly punctured than disk; sagittal line distinctly but finely impressed, almost reaching anterior margin, but oblitered towards base, because of longitudinal basal impression, excavated near each margin, open to lateral gutter; each point with a seta, quite long, inclined posteriorly. Elytra almost twice wider than pronotum, ovoid, longer by half than wide, quite obtuse near basal margins, quite distinctly descending from peduncle in flat line towards humerus, which however marked, is always very rounded; lateral margins quite regular from humeri towards apex, quite sinuate before apex, which weakly obtuse; dorsum regularly convex (distinctly less than in C. bifasciatus Fabr. = tomentosus Dejean ), striae quite deeply impressed, quite grossly and not densely punctured, punctuation in narrow line, not ranging intervals; these convex, densely covered by setae, setal cover more dense on internal intervals; first stria diverged near base and fall in the second; rudimentary prescutellum quite long, along scutellum and basal part of suture; IX interval with a continual rank of umbilic points, which distributed somewhat irregularly, more at the middle than near margins; lateral rim wide, elevated longwise; prolonged near base to peduncle, quite trenchant. Head beneath smooth; pronotal epipleura wide, almost smooth, excavated in a gutter, ist bottom more similar to episterma than external margin; prosternum throughout grossly punctured and covered by long setae, two other parts of sternum punctured and covered by setae, episterna punctured as well as prosternum. Venter punctured and quite densely covered by setae, with punctuation gross, similar to that on episterna, diffused on two anterior segments, and near margins of the two rest segments, where it is rare; elytral epipleura punctured and weakly covered by setae. Fully black, glossily, terminal parts of palpar articles and setae, covering legs, ferruginous; each elytron with two large orange-yellowish maculae as in C. nobilis (Boheman, 1848) ; humeral macula reaching from IX to more or less II stria [III interval], creates a wide fascia, sligthly serrate, weakly oblique and arcuate; preapical macula reaching from VIII to II stria [III interval], creates a fascia, little obliqued posteriorly to lateral margin, compounds of six macular spots of eaqual length, except spot on VII interval, prolonged weakly to apex, creating a small dent; anterior macular margin is not distinctly serrate. I have three specimens from Senegal and Senegambia Portuguese (Boccandé); in one specimen both elytral maculae (however this genus never is varying in elytral colouration) are more red than in two others, which undoubtedly, is an effect of alcohol or other chemicals” [from French].

Distribution. Guinea, Guinea-Bissau, Senegal.

Figures

PLATE 3. Habitus of C. strachani group III (Scale bar: 10 mm). 18. C. tetrastigma tetrastigma (Laferté-Sénectere, 1851), lectotype male designated by Chaudoir, 1879; 19. C. tetrastigma tetrastigma (Laferté-Sénectere, 1851), paralectotype designated by Chaudoir, 1879, female; 20. C. tetrastigma morettoi n. ssp., HT (male); 21. C. tetrastigma morettoi n. ssp., PT female, western Ivory Coast; 22. C. tetrastigma morettoi n. ssp., PT male, Mali; 23. C. tetrastigma morettoi n. ssp., PT female, eastern Ivory Coast; 24. C. tetrastigma milzi (Duvivier, 1891) male, Central Africa; 25. C. tetrastigma milzi (Duvivier, 1891) female, DR Congo: Orientale (HT of C. strachani depressus Alluaud, 1929); 26. C. tetrastigma milzi (Duvivier, 1891) male, DR Congo: Orientale (PT of C. strachani depressus Alluaud, 1929); 25. C. tetrastigma milzi (Duvivier, 1891) female, DR Congo: Orientale.

PLATE 12 Craspedophorus strachani group I, aedeagus in frontal and right lateral views: 91. C. strachani strachani (Hope, 1842), Sierra Leone; 92. C. strachani grossus (Hope, 1842), Togo; 93; C. strachani monardi (Basilewsky, 1951), Cameroon, North; 94. C. strachani bamendanus new subspecies, Cameroon, Northwest; 95. C. latipennis Burgeon, 1930; Cameroon, Southwest; 96. C. tetrastigma tetrastigma (Laferté-Sénectere, 1851), Guinea; 97. C. tetrastigma morettoi new subspecies, western Ivory Coast; 98. C. tetrastigma morettoi new subspecies, eastern Ivory Coast; 99. C. tetrastigma milzi Duvivier, 1891, Orientale, DR Congo; 100. C. latemaculatus Alluaud, 1930, Cameroon, Central.