Tegenaria campestris, (C. L. KOCH, 1834)

Bolzern, Angelo, Burckhardt, Daniel & Hänggi, Ambros, 2013, Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 168 (4), pp. 723-848 : 789-790

TEGENARIA CAMPESTRIS (C. L. KOCH, 1834) STAT. REV.

(FIG. 26K, L, O, P)

Aranea decemguttata Martini & Goeze , in Lister (1778 (1792): 288) (nomen oblitum).

Aranea campestris C. L. Koch, 1834: 124 , pl. 20, male.

Tegenaria campestris: C. L. Koch, 1841: 34 , 35, figs 615, 616.

Philoica campestris: Simon, 1864: 202 .

Malthonica campestris: Guseinov et al., 2005: 164 . No type material available.

Other material examined

Austria (21 ♂, 8 ♀) ; Germany (9 ♂, 8 ♀) ; Hungary (1 ♂, 2 ♀) .

Description

Measurements: Male (N = 1): CL 3.68, CW 2.93, STL 1.90, STW 1.75, OL 4.02, OW 2.64. Leg I (3.52, 1.46, 3.12, 3.18, 2.00), II (3.41, 1.39, 2.59, 2.82, 1.59), III (3.12, 1.15, 2.20, 2.90, 1.42), IV (3.93, 1.35, 2.98, 3.84, 1.71). Pedipalp (1.59, 0.62, 0.51, 1.78), bulbL 1.49. Female (N = 1): CL 3.65, CW 2.79, STL 1.94, STW 1.77. Leg I (3.59, 1.41, 3.14, 3.02, 1.89), II (3.20, 1.36, 2.45, 2.71, 1.60), III (2.94, 1.16, 2.00, 2.55, 1.22), IV (3.94, 1.37, 3.03, 3.74, 1.71). Pedipalp (1.35, 0.62, 0.77, 1.56). EPL 0.40, EPW 0.65, ATL 0.26, ATW 0.38. Eyes: PME 0.18, PLE 0.19–0.21, AME 0.14–0.16, ALE 0.20–0.21. Eye distances: PME–PME 0.5–1 x PME, PME–AME 0.5–1 x PME, PME–PLE 0.5–1 x PME, PME–ALE 0.5–1 x PME, AME–AME <0.5 x AME, AME–ALE << 0.5 x AME. CLY1 1.5–2 x AME, CLY2 1 x ALE.

Male palp: RTA with three branches, ventral branch extensive, lobe-like with distinct ridge, distally moderately protruding, lateral branch spur-like and strongly sclerotized, dorsal branch broad and strongly protruding, distally with several points and ridges. Filiform embolus length about 2.5–3 x CB, originating at 7–8 o’clock position, distal tip at 4–5 o’clock position. Conductor in retrolateral view S-shaped, distal portion distinctly elongated, lateral margin completely folded. Terminal end bifid, ventral part short and pointed, dorsal part plate-like, rounded. Connection of conductor to tegulum moderately sclerotized. MA originating at 6 o’clock position, strongly protruding, distally with complex plate-like sclerite (short point and a longer hook-like ending). MA membranously connected to tegulum. Basal portion of tegulum visible and undulated.

Epigyne and vulva: Epigyne with distinct median plate, anteriomedially continuously connected to strongly sclerotized epigynal plate. Posterior sclerite absent. CO anteriorly of median plate, distinctly visible as gaps. Vulva consists of CBD, no distinct RC recognizable. First part (CD) of CBD less sclerotized and moderately convoluted, proximal part strongly convoluted and sclerotized. FD only represented by small, leaf-shaped appendages.

Other important characters: Cheliceral promargin with three, retromargin with three to four teeth. Colulus developed as trapezoidal plate with distal margin straight or indistinctly notched. Same pattern of distal spigots on PMS (in females) as described for the type species. PLS with distal segment shorter than basal segment. Tarsal trichobothria on cymbium and palp tarsus absent. Tarsal trichobothria six to eight. Small teeth on paired claws of leg I seven to ten. Leg spination: male palp (2–0–0–0, 2–0–0, 2–2– 0–0), female palp (2–0–0–0, 2–0–0, 2–2–0–0), leg femora (2–2–1–0, 2–2–2–0 or 2–3–2–0, 2–2–1–0 or 2–2–2–0 or 2–3–2–0, 1–1–1–0 or 1–1–2–0 or 2–1–1– 0), patellae (all 2–0–0), tibiae (0–0–0–1+2p or 0–0–0– 1p+1+1p or 0–0–0–2p or 0–0–0–3p or 2–0–0–1+2p, 0–2–0–1+2p or 0–2–0–2+1p or 2–2–0–1+2p or 2–2–0– 2+1p, 2–2–2–1+2p or 2–2–2–2+1p or 2–2–2–3p, 2–2– 2–1+2p or 2–2–2–2+1p or 2–2–2–3p), metatarsi (0–0– 0–1+2p+1 or 0–0–0–3p+1, 0–2–0–1p+1+2p+1 or 0–2– 0–3p+1, 0–3–2–3p+1 or 0–3–3–3p+1, 0–3–3–1+3p+1 or 0–3–3–3p+1 or 0–3–3–4p+1), tarsi (I & II 0, III: 0–0– 1–0, IV: 0–0–1–0 or 0–1–1–0).

Coloration: Margin of carapace narrowly darkened with three to four crescent-shaped spots, dorsally with two symmetrical longitudinal dark bands. Sternum with distinct pattern of pale median region and three pairs of symmetrical pale dots laterally, somewhat fused together. Chelicerae with distinct, darkened spots. Opisthosoma dark brownish, anteriomedially with yellowish band, continuing in broad chevrons posteriad. Legs annulated. ALS basally darkened, PLS basal segment darkened, distal segment pale.

Distribution

Reported from Central Europe in the west to Azerbaijan in the east ( Blick et al., 2004; Otto & Dietzold, 2006; van Helsdingen, 2011).

Discussion

Tegenaria campestris is part of a species-complex including Teg. montana, Teg. rilaensis , and the recently described Teg. bozhkovi ( Deltshev, 1993; Deltshev, 2008b). The morphological differences amongst the species are very small. Owing to the limited number of specimens examined of Teg. montana, Teg. rilaensis , and Teg. bozhkovi , we could not judge if these differences represent inter- or intraspecific variation.