Tegenaria regispyrrhi, BRIGNOLI, 1976

Bolzern, Angelo, Burckhardt, Daniel & Hänggi, Ambros, 2013, Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 168 (4), pp. 723-848 : 813-814

TEGENARIA REGISPYRRHI BRIGNOLI, 1976

( FIG. 24A–D )

Tegenaria regispyrrhi Brignoli, 1976b: 569–571 , figs 54, 56, 58

Types

Holotype. Greece: Trikala : Malakasi, ♂ ( MCSN, 544), 28.ix.1966, Brignoli.

Paratypes. Same data as for holotype, 2 ♀; Ioannina: Metsovon, 1 ♀ ( MCSN, 100), 22.vii.1971, Osella; ‘ 3 km après Karies: en direction de Elati’, 1 ♀ ( MHNG), 26.iv.1973, Mahnert .

Other material examined

Greece: Ioannina : ‘ Passo Katava’, 1 ♂ ( MSNB), 4.vi.1991, Giachino; ‘ E Ioannina, near E92’, 1 ♀ ( SMF), 2.iv.2006, Schönhofer; Kefallonia: 1 ♀ ( SMF, coll. Roewer, 5972); Trikala: ‘street E92 btw. Panagia and Metsovo’, 1 ♂, 1 ♀ ( SMF), 2.iv.2006, Schönhofer .

Description ( Teg. regispyrrhi sensu stricto)

Measurements: Male (N = 1): CL 2.84, CW 2.14, STL 1.45, STW 1.37. Leg I (2.87, 1.08, 2.67, 2.74, 1.81), II (2.74, 0.99, 2.27, 2.50, 1.62), III (2.59, 0.93, 1.98, 2.54, 1.38), IV (3.21, 1.03, 2.78, 3.46, 1.74). Pedipalp (1.58, 0.58, 0.56, 1.35), bulbL 0.98. Female (N = 2): CL 2.11–

3.20, CW 1.54–2.22, STL 1.17–1.59, STW 1.08–1.45. Leg I (2.07–2.75, 0.83–1.01, 1.80–2.41, 1.75–2.47, 1.24–1.48), II (1.92–2.67, 0.76–1.08, 1.49–2.11, 1.61– 2.34, 1.02–1.45), III (1.82–2.44, 0.68–1.01, 1.31–1.89, 1.57–2.35, 0.97–1.26), IV (2.23–3.22, 0.71–1.11, 1.80– 2.78, 2.28–3.31, 1.13–1.51). Pedipalp (0.87–1.30, 0.41– 0.56, 0.58–0.78, 0.88–1.21). EPL 0.32–0.45, EPW 0.52– 0.70, ATL 0.14–0.25, ATW 0.36–0.51. Eyes: PME 0.11– 0.14, PLE 0.11–0.17, AME 0.07–0.09, ALE 0.12–0.14. Eye distances: PME–PME 0.5–1 x PME, PME–AME 0.5–1 x PME, PME–PLE ≤ 0.5 x PME, PME–ALE 0.5–1 x PME, AME–AME ≤ 0.5 x AME, AME–ALE << 0.5 x AME. CLY1 2.5–3 x AME, CLY2 1 x ALE.

Male palp: RTA with three branches, ventral branch indistinct, flat, long, drawn-out bulge with distinct ventral ridge, lateral branch strong and straight, protruding, thorn-like, dorsal branch large, leaf-shaped, and strongly protruding, bent and distally pointed. Filiform embolus length about 1.5 x CB, originating at 8–9 o’clock position, distal tip at 4 o’clock position. Conductor irregularly shaped with distal portion strongly elongated and moderately bent, lateral margin completely folded. Terminal end not bifid, indistinctly pointed. Connection of conductor to tegulum membranous. MA originating at 5–6 o’clock position, strongly protruding, distally with bifid platelike sclerite, one part hook-like and elongated, the other part broadly pointed. MA membranously connected to tegulum. Basal part of tegulum clearly visible, moderately undulated.

Epigyne and vulva: Epigyne with distinct atrium (even though the CO are not situated in this atrium), anteriorly separated from the epigynal plate by a prominent triangularly or trapezoidally shaped ridge. Posterior sclerite absent. CO anterolateral of the prominent ridge distinctly visible as gaps. Epigynal ‘pseudo teeth’ absent. Vulva consists of distinguishable CD, RC, and FD, all structures are fused together into a strongly sclerotized structure. CD long and straight (difficult to observe), leading into globular RC, RC separated by about their diameter. FD long and convoluted, terminally leading into small, leaf-shaped appendages.

Other important characters: Cheliceral promargin with three, retromargin with three to four teeth. Colulus developed as trapezoidal plate with the distal margin medially notched. Same pattern of distal spigots on PMS (in females) as described for the type species. PLS with distal segment shorter than basal segment. Tarsal trichobothria on cymbium and palp tarsus absent. Tarsal trichobothria five to seven. Small teeth on paired claws of leg I nine to ten. Leg spination: male palp [2–0–0–0, 2–0–0, 0–2–0–0 (not clear if these are dorsal or prolateral spines)], female palp (2–0–0–0, 2–0–0, 2–1–1–0 or 2–2–0–0), leg femora (2–2–0–0 or 2–2–1–0, 2–1–2–0 or 2–2–0–0, 2–1–1–0 or 2–2–2–0, 1–1–1–0 or 2–1–1–0 or 2–2–1– 0), patellae (all 2–0–0), tibiae (0 or 2–0–0–1+1p or 2–0–0–3p or 2–1–0–3p or 2–2–0–3p, 2–1–0–2+1p or 2–1–0–3p or 2–2–0–1+1p or 2–2–0–2+1p, 2–2–2–2+1p or 2–2–2–3p, 2–2–2–2+1p or 2–2–2–3p), metatarsi (0–0–0–3p+1, 0–1–0–3p+1, 0–3–3–3p+1 or 1–3–3– 3p+1, 1–3–3–3p+1), tarsi (I & II 0, III & IV 0–0–1–0).

Coloration: Margin of carapace with three narrow, crescent-shaped, darkened spots, sometimes connected, dorsally with two symmetrical longitudinal dark bands, serrated. Sternum with distinct pale median band and three symmetrical pairs of pale spots laterally. Opisthosoma dark brownish, laterally with yellowish spots, dorsally with indistinct yellowish chevrons. Legs annulated. Colulus partly darkened, ALS darkened, PLS with basal segment darkened (pale in other specimens, may be a result of alcohol preservation), distal segment pale.

Distribution

Reported from Greece.

Figures

Figure 24. A–D, Tegenaria regispyrrhi; E, F, Tegenaria aff. regispyrrhi 1; G, H, Tegenaria aff. regispyrrhi 2; I, J, Tegenaria aff. regispyrrhi 3; K-P, Tegenaria vankeerorum sp. nov. Left male palp in ventral (C, K) and retrolateral views (D, L); retrolateral tibial apophysis of left male palp in dorsal view (M); female epigyne in ventral (A, E, G, I) and vulva in dorsal view (B, F, H, J); face in frontal view (N); sternum (O) and spinnerets (P) in ventral view. Scale bars = 0.5 mm.