Filago abyssinica Schultz Bipontinus ex Richard (1848: 432)

Andrés-Sánchez, Santiago, Martínez-Ortega, M. Montserrat & Rico, Enrique, 2016, Typifications and a new name in Filago (Gnaphalieae, Asteraceae), Phytotaxa 243 (3), pp. 281-290 : 285-287

Filago abyssinica Schultz Bipontinus ex Richard (1848: 432)

Lectotype (designated here): ― ETHIOPIA. In agriis frumentariis prope Adoua, 10 October 1842, Schimper 1913 ( G00223822 photo!).

Additional specimens examined: ― ETHIOPIA. In agriis frumentariis prope Adoua, 10 October 1842, Schimper 1913 (isolectotypes G00223821 photo!; G00223823 photo!; G00223824 photo!; K274251 !; MPU!; GOET001546 !; TUB005090 photo!) .

Protologue citation: ― “In agriis frumentariis prope Adoua (Schimper)”.

Note: ― Wagenitz (1969) indicated three sheets preserved at G, GOET and TUB as original material for F. abyssinica but he did not choose a lectotype. We select the herbarium specimen kept at Genève (G00223822) as the lectotype and indicate five isolectotypes.

Filago apiculata G.E.Sm. ex Babington (1851: 174) , nom. illeg. (Art. 52.1 ICN) = Filago lutescens Jordan (1846: 201)

Lectotype (designated here): ― UNITED KINGDON. Cantley, August 1845, G. E. Smith s.n. ( K372682 !).

Additional specimens examined: ― UNITED KINGDON. Cantley, Yorkshire, G. E. Smith s.n. (syntypes K372680 !; K372681 !) ; Cantley near Doncaster, G. E. Smith s.n. (syntype BM! Mrs Robinson’s Herbarium) .

Protologue citation: ― “Sandy borders of fields, hedge-banks and road-sides, Cantley, Rossington, & c. near Doncaster”.

Filago arvensis var. gracilis Sennen (1929: 39) = Filago arvensis Linnaeus (1753: 856)

Lectotype (designated here): ― SPAIN. Cerdagne, Val de l’Estahuja , sables du ruisseau, 1250 m, 25 July 1920, F. Sennen 3957 ( BC874213 ! mounted on two sheets).

Additional specimens examined: ― SPAIN. Cerdagne, Val de l’Estahuja , sables du ruisseau, 1250 m, 25 July 1920, F. Sennen 3957 (isolectotypes SEV212946 !; MPU! MPU-COSTE) .

Protologue citation: ― “Vall de l’Estahuja, par les alluvions du ruiseau, vers 1250 m ”.

Filago arvensis var. ramosa Rouy (1903: 175) = Filago arvensis Linnaeus (1753: 856)

Lectotype (designated here): ― Illustration in Oeder, Fl. Dan. Tabula MCCLXXV, 1806.

Epitype (designated here): ― SPAIN. Ávila, Piedrahita, Santiago del Collado, Arroyo del Prado de los Peces, 1593 m, 21 June 2006, Andrés-Sánchez, García-Barriuso, González-Canalejo & Rico SA85 (SALA134283!).

Protologue citation: ― “Fries Nov. Ed. 2, p. 267; Fl. Dan., t. 1275”.

Note: ― The illustration chosen as lectotype is the only original material found for this name. But it is not depicted with enough resolution and some morphological details (e.g., indumentum of the achenes, and consistence of the involucral bracts) are not observable. These characters are essential to discriminate this taxon from closely related species. For these reasons we propose an epitype, which is one of the samples included in the phylogenetic analyses of DNA sequence data presented in Galbany-Casals et al. (2010).

Filago arvensis var. subsimplex Rouy (1903: 175) = Filago arvensis Linnaeus (1753: 856)

Lectotype (designated here): ― SWEDEN. Upsaliae in montosis, August 1840, Fries s.n. ( E. Fries, Herbarium Normale, Fasc. VII, Nº 2) ( P! Herb. Schultz Bip., Herb. E. Cosson).

Additional specimens examined: ― SWEDEN. Upsaliae in montosis, August 1840, Fries s.n. ( E. Fries, Herbarium Normale, Fasc. VII, Nº 2) (isolectotypes P! Herbier E. Drake; P!) ; Upsaliae, August, Fries s.n. ( E. Fries, Herbarium Normale, Fasc. VII, Nº 2) (isolectotype P! Herb. Schultz Bip., Herb. E. Cosson) ; in jugis arenosis juxta Upsaliam, 25 July 1861, Jelterstedt Reliquiae Mailleanae 1307 (syntypes P! Herbier E. Drake; P! Donné par la famille de M. Maille; P! Herbier Th. Delacour; MPU! Herbier du Dr. Le Sourd ; MPU! Herbier Guillon; K! Herb. F. A. Rogers) ; FRANCE. Sables arides à Haguenau, C. Billot 38 (syntypes P! Herbier E. Drake; MPU! Herbier Bernardine Martin; MPU!; MPU! Herbier du Dr. Le Sourd ; MPU! Herbier de la Faculté de Médecine; MPU! MPU-ASCNRS; P!; P! exsiccata Th. Delacour; P!) .

Protologue citation: ― “Fries Herb. Norm., 7, nº 2; Reliq Maill., 1307; Bill., 38”.

Filago atlantica Ball (1873: 364) = Filago discolor ( Candolle 1836: 459) Andrés-Sánchez & Galbany in Galbany-Casals et al. (2010: 1689)

Lectotype (designated here): ― MOROCCO. Ex regiones subalpina Atlantis Majoris in convalle Ait Mesan, supra Arround, 13–15 May 1871, J. Ball s.n. ( K274236 !).

Additional specimens examined: ― MOROCCO. Ex regiones subalpina Atlantis Majoris in convalle Ait Mesan, supra Arround, 13–15 May 1871, J. Ball s.n. (isolectotype P! Herbier E. Cosson).

Protologue citation: ― “In regione superiori Atlantis Majoris. In convalle Ait Mesan supra Arround! A 2200m ad 2400m in glareosis arenaceis”.

Filago bianorii Sennen & Pau in Sennen (1913: 42) = Filago congesta Gussone ex Candolle (1838: 248)

Lectotype (designated here): ― SPAIN. Baleares: Sables maritimes de Molinar à Palma, July 1911, F. Bianor 1261 ( BC874215 !).

Additional specimens examined: ― SPAIN. Baleares: Sables maritimes de Molinar à Palma, July 1911, F. Bianor 1261 (isolectotypes LD1019588 photo!; MA124165 !; MPU-COSTE!; M) .

Protologue citation: ― “ Baléares: sables maritimes du Molinar, près Palma. ( F. Bianor.)”.

Note: ― Wagenitz (1966) indicated two vouchers preserved at LD and M as original material for F. bianorii but he did not choose a lectotype. We have found three additional sheets, which are original material for this name. We propose BC874215 as lectotype.

Filago bolivarii Caballero (1915: 287) = Filago congesta Gussone ex Candolle (1838: 248)

Neotype (designated here): ― MOROCCO. Mont-Aroui , Tiztoutine, 34º59’43.3’’N 3º3’25.7’’W, 14 March 2008, Andrés-Sánchez, Martínez-Ortega, Peñas & Lorite SA147 ( SALA134198 ).

Protologue citation: ― “Habitat in cultis prope Melilla in loco dicto Posada del Cabo Moreno. April”.

Note: ― There is a voucher kept at MA (MA124064!) containing plants collected by Caballero in Posada del Cabo Moreno in April 1912. This sheet could be considered as possible original material relevant for the typification of the name F. bolivarii . But it consists of three individuals, which clearly correspond to F. pyramidata Linnaeus (1753: 1199) . However, Caballero in the protologue was clearly trying to describe F. congesta , a species characterized by bearing inner florets functionally male with ovary sterile (“floribus disci 5–6 ebracteatis sterilibus”) and by its achenes lacking a pappus (“achaeniis disci eppaposis”). The sheet MA124064 contains specimens that show inner florets hermaphrodite with fertile ovaries and achenes with pappus. These characters are so evident and in apparent conflict with the protologue, that it seems reasonable to accept that they were not used to elaborate the validating description of F. bolivarii . As no further possible original material has been found, we choose a neotype for this name, which is in accordance with the protologue. We select as neotype a collection from a locality close to that of the type.

Filago canescens Jordan (1846: 202) = Filago germanica ( Linnaeus 1753: 857) Hudson (1762: 328)

Lectotype (designated here): ― FRANCE. Lyon , 1845, Jordan s.n. ( LYJB Jordan VII-A11 photo!).

Additional original material examined: ― FRANCE. De Lyon culta, Jordan s.n. (syntype LYJB Sagot VI-H7 photo!) ; Illustration in Jordan , Observ. Pl. Nouv. 3: pl. 7, f. A, 1–10, 1846 .

Protologue citation: ― “…aux environs de Lyon…”.

Filago cuneata Lojacono (1903: 107) = Filago fuscescens Pomel (1874: 44)

Lectotype (designated here): ― ITALY. La Favorita, May 1850 ( PAL008596 photo!).

Additional specimens examined: ― ITALY. La Favorita (isolectotypes PAL008597 photo!; PAL008598 photo!; PAL008600 photo!; PAL008604 photo!) .

Protologue citation: ― “in pascuis secchi marittimi alla Favorita, abbevaratojo delle Lenti Herb. Pan. Manu Tin.! Marsala Sciare Herb. Pan.!”.