Ellimenistes humeralis, Marshall, R. Borovec, 1947

Borovec, Roman & Nakládal, Oto, 2019, Ellimenistes humeralis Marshall, 1947 and its allies (Coleoptera: Curculionidae: Entiminae: Embrithini), Zootaxa 4563 (1), pp. 175-184 : 183

Key to the species of the Ellimenistes humeralis group

1. Rostrum narrowest at base ( Fig. 4 ). Elytra with laterally prominent projections behind the base ( Fig. 3 ). Funicle segments 1 and 2 equally long. Eyes big, conspicuously prominent from outline of head ( Fig. 4 ). Scrobes laterally furrow-shaped ( Fig. 7 ). Pronotum 1.6–1.8× wider than long, widest at base, base V-shaped ( Fig. 3 ). Raised elytral setae at apical declivity longer than at basal half. Ramus of spermatheca shorter than wide ( Figs 16, 18 ).............................................. 2

- Rostrum widest at base ( Fig. 5 ). Elytra without laterally prominent projections behind the base ( Fig. 2 ). Funicle segment 1 twice longer than segment 2. Eyes small, barely prominent from outline of head ( Fig. 5 ). Scrobes laterally subtriangular ( Fig. 8 ). Pronotum 1.3–1.4× wider than long, widest at basal third, base straight ( Fig. 2 ). Raised elytral setae equally long at basal and apical part. Ramus of spermatheca twice longer than wide ( Figs 17, 19 )....................................... 3

2. Setae at posterior elytral declivity appressed, as short as one third of width of an elytral interval ( Fig. 1 ). Elytra with rounded sides ( Fig. 1 ). Appressed elytral scales isolated. Spermatheca with nodulus about 6× longer than wide, U-shaped curved ( Fig. 16 )................................................................................ E. humeralis Marshall

- Setae at posterior declivity erect, as long as width of one interval ( Fig. 3 ). Elytra with almost straight sides ( Fig. 3 ). Appressed elytral scales slightly imbricate. Spermatheca with nodulus twice longer than wide, perpendicularly curved ( Fig. 18 )............................................................................................ E. marshalli sp. nov.

3. Rostrum shorter, 1.41–1.44× wider than long ( Fig. 5 ), in profile distance between scrobe and eye equally long as scrobe ( Fig. 8 ). Funicle segment 1 1.8–2.0× longer than segment 2; segments 3–7 isodiametric. Penis widest at base, with sides before apex slightly convex ( Fig. 10 ). Tegmen with parameres distinctly shorter than diameter of ring, separated ( Fig. 13 ). Plate of female sternite VIII widely subtriangular, apically constricted ( Fig. 21 ). Spermatheca with ramus pressed to corpus ( Fig. 17 ).............................................................................................. E. janaki sp. nov.

- Rostrum longer, 1.23–1.28× wider than long ( Fig. 6 ), in profile distance between scrobe and eye longer than scrobe ( Fig. 9 ). Funicle segment 1 1.3–1.4× longer than segment 2; segments 3–7 1.1× longer than wide. Penis widest at apical quarter with sides before apex slightly concave ( Fig. 12 ). Tegment with parameres almost as long as diameter of ring, connected at their basal half ( Fig. 15 ). Plate of female sternite VIII narrowly subtriangular, without apical constriction ( Fig. 23 ). Spermatheca with ramus deviated from corpus ( Fig. 19 ).................................................... E. raucus sp. nov.

Figures

FIGURES 4–19. Structural details of Ellimenistes Boheman. Figs 4–6, head with rostrum, dorsal view: 4. E. humeralis Marshall; 5. E. janaki sp. n.; 6. E. raucus sp. n. Figs 7–9, head with rostrum, lateral view: 7. E. humeralis Marshall; 8. E. janaki sp. n.; 9. E. raucus sp. n. Figs 10–12, penis, ventral and lateral view: 10. E. janaki sp. n.; 11. E. marshalli sp. n.; 12. E. raucus sp. n. Figs 13–15, tegmen, dorsal view: 13. E. janaki sp. n.; 14. E. marshalli sp. n.; 15. E. raucus sp. n. Figs 16–19, spermatheca: 16. E. humeralis Marshall; 17. E. janaki sp. n.; 18. E. marshalli sp. n.; 19. E. raucus sp. n.

FIGURES 1–3. Dorsal habitus of Ellimenistes Boheman. 1. E. humeralis Marshall; 2. E. janaki sp. n.; 3. E. marshalli sp. n.

FIGURES 20–29. Structural details of Ellimenistes Boheman, Cycliscus Schoenherr and Cosmorhinus Schoenherr. Figs 20– 25, sternite VIII of female, dorsal view: 20. Ellimenistes humeralis Marshall; 21. E. janaki sp. n.; 22. E. marshalli sp. n.; 23. E. raucus sp. n.; 24. Cycliscus caecicollis Boheman; 25. Cosmorhinus cervinus Boheman. Figs 26–29, ovipositor, dorsal view: 26. Ellimenistes humeralis Marshall; 27. E. janaki sp. n.; 28. E. marshalli sp. n.; 29. E. raucus sp. n.