Nannaria sigmoidea (Hennen & Shelley, 2015)

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

publication ID

http://dx.doi.org/10.3897/zookeys.1030.62544

publication LSID

lsid:zoobank.org:pub:48997D3B-9B4E-4348-8F30-0C8E8672569A

persistent identifier

http://treatment.plazi.org/id/BBAFCC7B-1C0D-525B-B168-0C002B8A6BBF

treatment provided by

ZooKeys by Pensoft

scientific name

Nannaria sigmoidea (Hennen & Shelley, 2015)
status

 

Nannaria sigmoidea (Hennen & Shelley, 2015)   Vernacular name: "The Sigmoid Gonopod Twisted-Claw Millipede" Fig. 112 View Figure 112

Mimuloria dilatata sigmoidea   Hennen & Shelley, 2015: 1-15, figs 20, 21.

Nannaria sigmoidea   : Means et al. 2021: S72.

Material examined.

Holotype: United States - Tennessee • ♂; Meigs County, c.a. 7.2 km S Decatur, along country road 4274, 0.8 km from junction of Tennessee highway 58; [35.4378°N, - 84.9140°W]; 14 Oct. 1978; R. M. Shelley, W. B. Jones leg.; NCSM NCSM27946. For detailed collection data see Suppl. material 7. GoogleMaps  

Diagnosis.

Adult males of N. sigmoidea   are distinct from other Nannaria   , including the nearby N. blackmountainensis   sp. nov. and Nannaria   sp. nov. ‘Cratagae’ ( Nannaria wilsoni   species group), based on the following combination of characters: Gonopods. Gonopodal acropodite simple and gently curving anteromedially, not straight as in Nannaria   sp. nov. ‘Cratagae’ or with medial swelling as in N. blackmountainensis   sp. nov. Acropodite with prominent, rounded lateral flange near tip (Fig. 112A View Figure 112 , red arrow), not hooked as in N. blackmountainensis   sp. nov. or lacking as in Nannaria   sp. nov. ‘Cratagae.’ Acropodite tip directed posteriorly, not dorsally as in N. blackmountainensis   sp. nov. Telopodite basal zone subequal to length of prefemoral process, not ca. 1/2 length as in N. blackmountainensis   sp. nov. or ca. 1/3 length as in Nannaria   sp. nov. ‘Cratagae.’ Prefemur with prefemoral process bent at 90° angle at halfway point, directed medioventrally, not straight, acicular as in N. blackmountainensis   sp. nov. and Nannaria   sp. nov. ‘Cratagae.’ Prefemoral spine reduced to small, blunt lobe at base of prefemoral process (Fig. 112A View Figure 112 , red triangle), not fused as a small ridge as in N. blackmountainensis   sp. nov. or lacking as in Nannaria   sp. nov. ‘Cratagae.’ Color. Color in life unknown.

Measurements.

♂ holotype ( NCSM, NCSM27946): BL = 36.1, CW = 5.0, IW = 3.0, ISW = 1.1, B11W = 6.0, B11H = 3.8.

Variation.

No known variation.

Distribution.

Known only from the type locality (Tennessee: Meigs County; Suppl. material 7; Fig. 129 View Figure 129 ). Distribution area: N/A; status: MRE.

Ecology.

Hennen and Shelley (2015) note that only that the type specimens were collected from the east side of the Tennessee River.

Etymology.

Hennen and Shelley (2015: 15) stated that "[t]he subspecific name denotes the overall sigmoidal curvature of the prefemoral process, which also passes through numerous vertical planes."

Note on clade membership.

No specimens of Nannaria sigmoidea   were collected for this study. Nannaria sigmoidea   shares gonopodal characters with members of the Nannaria ambulatrix   clade, such as a prominent lateral flange on the acropodite tip, a prefemoral process bent at 90°, and crossing of the acropodites in the medial view (found in N. ambulatrix   sp. nov. and N. botrydium   sp. nov.). However, due to the wide geographic separation between N. sigmoidea   and the Nannaria ambulatrix   clade (> 270 km), and the presence of these characters in a member of the Nannaria minor   clade ( N. kassoni   sp. nov.), the phylogenetic placement of N. sigmoidea   in a clade is uncertain.

Type locality.

United States, Tennessee, Meigs County, c.a. 7.2 km S Decatur, along country road 4274, 0.8 km from junction of Tennessee highway 58.

Notes.

In the original publication, Hennen and Shelley (2015: 15) designate a male holotype collected by R. M. Shelley and W. B. Jones on October 14, 1978.