Lasiacis maculata (Aubl.) Urb.

Baldini, Riccardo M., 2010, Panicum maculatum Aubl.: its type and the correct use of Lasiacis maculata (Aubl.) Urb. (Poaceae, Paniceae), Adansonia (3) 32 (1), pp. 109-119 : 114-118

Lasiacis maculata (Aubl.) Urb.

Symbolae Antillanae 8: 751. (1921). — Panicum maculatum Aubl., Histoire des Plantes de La Guiane Françoise 1: 51 (1775). — Type: (unpublished iconography from a plant of Martinique) “ Milium arundinaceum scandens et maculatum ” in Plumier’s Botanicon americanum MS 4!: t. 82 (holo-, Bibliothèque centrale du Muséum national d’Histoire naturelle!) ( Figs 1 ; 2 ). — Martinique, environs de St. Pierre, Mai 1853, Belanger 390 (epi-, FI-Webb! [ Fig. 3 ], here designated; isoepi-, G!, P! see also http://www.mnhn.fr/base/ sonnerat.html).

Panicum lanatum Sw. , Prodromus descriptionum Vegetabilium maximam partem incognitorum quae sub Itinere in Indiam Occidentalem 24. 1788, non Rottb. (1776), nom. illeg. — Type: Jamaica, Swartz s.n., (holo-, S; iso-, BM!).

Panicum glutinosum Lam., Tableau encyclopédique et méthodique 1: 174, tab. 43, fig. 3 (1791), non Sw. (1788), nom. illeg. — Panicum agglutinans Kunth, Enumeratio Plantarum 1: 120. (1833), nom. illeg. based on P.glutinosum Lam. ( Lamarck 1791) . — Panicum divaricatum L. var. agglutinans (Kunth) Hack. ex Sodiro, Anales de la Universidad Central del Ecuador 3 (25): 478 (1889). — Type: Tab. 43, fig. 3. in Lamarck (1791) (lecto-, here designated).

Panicum sorghoideum Desv. in Ham., Prodromus Plantarum Indiae Occidentalis 10 (1825). — Panicum lanatum Sw. var. sorghoideum (Desv.) Griseb. , Flora of the British West Indian Islands 551 (1864). — Lasiacis sorghoidea (Desv. in Ham.) Hitchc. & Chase, Contributions from the United States National Herbarium 18: 338 (1917). — Type: Porto Rico, Desvaux Herb. 24. (holo-, P!).

Panicum orinocense Willd. ex Spreng. , Systema Vegeabilium 1: 316 (1825), pro syn. — Type: Humboldt s.n., P. orinocense 1, Venezuela, Orinoco, s.d., (lecto-, B-W!, here designated; isolecto-, B-W! [5 sheets!], BM!).

Panicum arborescens Sieb. ex Trin., De Graminibus paniceis 208. 1826, non L. (1753), pro syn. — Type: Martinique, Sieber 267 (lecto-, BM!, here designated; isolecto-, FI-Webb!, GOET!, HAL!, L!, P!, W!).

Panicum divaricatum L. var. latifolium Schltdl. & Cham. , Linnaea 6: 33 (1831). — Type: Sieber 120 (lecto-, HAL!; isolecto, L!).

Panicum divaricatum L. var. lanatum Schltdl. & Cham. , Linnaea 6: 33. (1831). — Type: Jamaica, Willdenow Herb. 18760 (holo-, B).

Panicum fuscum Sieb. ex C.Presl, Abhandlungen der Böhmischen Gesellschaft der Wissenschaften , nebst der Geschichte derselben 3: 550 (1845), pro syn. —- Panicum martinicense Griseb. , Flora of the British West Indian Islands 552. 1864, nom. illeg., based on P. fuscum Sieb. ex C.Presl (1845) . — Panicum fuscum Sieb. ex Griseb. , Flora of the British West Indian Islands 552. 1864, pro syn. — Type: Martinique, Sieber, 1822, Flora Martin. No. 29 (lecto-, GOET! designated by Hitchcock [1920]; isolecto-, FI-Webb!, HAL!, L!, MO, P!, US, W!).

Panicum praegnans Steud. , Synopsis Plantarum Glumacearum 1: 74 (1854). — Panicum maculatum Aubl. var. pilosum Fourn., Mexicanas plantas 32 (1886), nom. illeg., based on P. praegnans Steud. — Type: Mexico, Oaxaca Buchinger 488 (lecto-, P! designated by Hitchcock [1920]).

Panicum guaraniticum Speg., Anales de Sociedad Científica Argentina 16: 107 (1883). — Lasiacis guaranitica (Speg.) Parodi, Notas del Museo de La Plata, Botánica 8: 95 (1911). — Type: Argentina, Misiones, Posadas , Mar 1883, von Gulich s.n. (holo-, LPS).

Panicum swartzianum Hitchc., Contributions from the United States National Herbarium 12: 140 (1908), nom. illeg., based on Panicum lanatum Sw. — Lasiacis swartziana (Hitchc.) Hitchc. , Botanical Gazette (Crawfordsville) 51: 302 (1911).

Lasiacis patentiflora Hitchc. & Chase, Contributions

from the United States National Herbarium 18: 338

(1917). — Lasiacis sorghoidea (Desv.) Hitchc. & Chase

var. patentiflora (Hitchc. & Chase) Davidse, Annals

of the Missouri Botanical Garden 64: 375 (1978). —

Type: Tobago, 20.XII.1912, Hitchcock 10268, (holo-,

US; iso-, US, L!, W!). Lasiacis acuminata Swallen, Memoirs of the New York

Botanical Garden 9: 267 (1957). — Type: Venezuela,

Territorio Amazonas, Great Rapids of the Orinoco,

Isla Carestia at Saltos Carestia y Galio, 5 km north

of Sanariapo, 100-200 m, erect to 2 m, 11.XI.1953,

Maguire, Wurdack & Bunting 36171 (holo-, US;

iso-, NY!). REPRESENTATIVE SELECTED MATERIAL SEEN FROM WEST INDIES AND GUYANAS Lasiacis maculata (Aubl.) Urb. (previously named under L. sorghoidea (Desv. in Ham.) Hitchc. & Chase). Puerto Rico. S.l., 1827, Wydler 222 (FI-Webb). — Maricao, 13.XI.1884, Sintenis 215 (L). — Coamo, 13.XI.1885, Sintenis 3062 (K, P, W). — Aybonito, 25.XI.1885, Sintenis 2861 (K). — Two miles NE of Mayaguez, 23.I.1900, Heller 4375 (L). — Vicinity of San Juan, 6.XI.1913, Chase 6412 (L). — Rosario Road, Monte Mesa, 12.XII.1913, Chase 1809 (L, W). — Vicinity of Mayagüez, Monte Mesa, 12.XII.1913, Chase 6824 (L). Virgin Islands. Scenic Drive, W slope of Mt. Eagle, 12.I.1980, Fosberg 5922 (BM). Antigua. 15.X.1931, Box 115 (BM, K). St. Kitts. 28.I.1932, Box 160 (BM, NY, K). Martinique. 1868, Husnot 94 (P). — Vallée de St. Pierre, V.1870, Hahn 105 (K, P). Grenada. 6.XII.1889, Eggers 6224 (P). — Bequia, 1950, Howard 11276 (BM). — St. Patrick, Marli Hill, 1.XII.1957, Proctor 17293 (BM). Dominica. Vicinity of Sophia Bay, 21.I.1966, Chambers 2608 (BM). — St. David near Madjini, 2.II.1969, De Filipps 191 (BM). Guadeloupe. S.d., Bertero s.n. (FI-Webb, TO). — 1839, Beaupertuis s.n. (P). — Gourbeyre, 31.I.1934, Rodriguez 2589 (P). — Grands Fonds de Saint-Anne, 18.XII.1987, Fournet 4272 (P). — Petit Bourg, Montebello, 16.IV. 1989, Fournet 4482 (P). Trinidad. Port of Spain, Fort Gorge Road, edge of jungle, 27.XI.1912, Hitchcock 592 (BM, C, K, L, W). — St. Anns near Port of Spain, 28.XI.1912, Hitchcock 594 (BM, C, K, L, P, W). — Trinidad, 19.XI.1913, Broadway 4841 (K). — St. Ann, 17.III.1920, Britton & al. 665 (K). — Aripo Road, 15.I.1926, Broadway 5991 (K). — Maraval, 9.XII.1928, Broadway 706 (BM, K). — Maracas, 26.VI.1929, Broadway 7233 (BM, G). — St. Anns, 21.II.1933, Broadway 9064 (BM, G). — Arima Valley, 1961, Snow s.n. (K). — Guanaco River 4 miles N de Maturita près Arima, 29.V.1975, Raynal 15598 (K). Tobago. Pigeon Hill, 7.IV.1959, Cowan 1448 (K, NY).— Parlatuvier-Roxborough Trace, Bloody Bay, 8.IV.1959, Cowan 1484 (K, NY, P). — Between Studley Park and Hillsborough Dam, 24.III.1993, Clement & Ryves 270 (BM).

Guyana. Oreala , Corentyne River, X.1879, Jenman 256 ( K) . — Rupununi , western extremity of Kanuku Mts., 4-22.III.1938, Smith 3314 ( K, P, W) . — Rupununi , Kanuku Mts., 14.XI.1957, Cook 173 ( K) . — Rupununi , 1 mile S of Emprensa, 14.IX.1963, Goodland 715 ( K, NY, P) . — Kanuku Mts. , slope of Nappi Mt., 8.XI.1987, Jansen-Jacobs & al. 758 ( K) . — Kanuku Mts. , Maipama, 17.XI.1987, Jansen-Jacobs & al. 966 ( K) . — Rupununi Distr. , Karanambo, 1.IX.1988, Maas & al. 7150 ( K) . — Rupununi , Eastern Kanuku Mts., NE of Warimure, 23. I.1991, Jansen-Jacobs 2182 ( K, NY, P) ; Rupununi , Kanuku Mts., Crabwood Creek, 25. I.1994, Jansen-Jacob & al. 3346 ( K, P) . — Rupununi , NE slope of Mount Shiriri, 18. VI.1995, Jansen-Jacobs & al. 4149 ( K, NY) .

French Guiana. S.l., 1842, Melinon 397 ( L, P). — Route de Kourou, 1.IX.1955 , Hoock 83 ( NY, P). — Route de Kourou, 28.IX.1955 , Hoock s.n. ( K, NY, P). — Saül , 30. VI.1956, Hoock 91 ( P). — Mount Saint-Marcel , 27. VI.2002, De Granville & al. 15518 ( K).

Suriname. Voltzberg, 21.VIII.1920, Pulle 227 ( L) . — Granitic plateau ± 8 km in line from Paka Paka (Saramacca R.) to Ebbatop (v. Asch v. Wijck-Range), 19.II.1951, Florschütz 1563 ( C) . — Circa portum aeronauticum ad flumen Oelemari, 1.IV.1963, Wessels Boer 1158 ( C) . — Lely Mts., 21.XI.1975, Lindeman & al. 158 ( K, NY, P) .

Lasiacis ligulata Hitchc. & Chase

Puerto Rico. 1827, Wydler 250 (FI-Webb, G) . — Puerto Rico , XII.1883, Stahl 7 ( W) . — Maricao, 13.XI.1884, Sintenis 215 ( BM, K, NY, W) . — Prope Lares in fruticetis ad “Palina Llanos”, 17. I.1887, Sintenis 5918 ( C) . — Jajome Alto , vicinity of Cayey, 3.XII.1913, Chase 6747 ( L) . — Near Maricao, Indiera Fria , 19-22.II.1915, Britton & al. 4490 ( K, NY) .

Dominica. S.coll. 344 ( K).

St. Kitts. S.d., Walsh s.n. ( K).

St. Thomas. 1802, Ventenat s.n. ( BM). — St. Thomas , 11.XII.1905, Raunkiaer 2304 ( C). — St. Thomas, Water Island, 3. I .1914, Ostenfeld 55 ( C).

St. Croix. Prosperity Garden, 5. I.1906, Raunkiaer 642 ( C).

Martinique. 1870, Hahn 1158 ( K).

St. Lucia. 6.XI.1929, Sandwith s.n. ( K)

Guadeloupe. S.d., Bertero s.n ( TO).

Trinidad. Green Hill , 2.XII.1910, Broadway 4038 ( BM) . — Trinidad, above St. Anns, near Port Spain , among bushes along strema, high climbing, 28.XI.1912, Hitchcock 10007 (= Am. Grass Natl. Herb. No. 589, holo-, US; iso-, BM!, G!, K!, NY!, P!, W!). — Tabaquite, 10.X.1912, Hitchcock 10170 ( L) . — Cedros , 12-14. XII.1912, Hitchcock 10157, ( L) . — Cedros , XII.1914, Hitchcock 33391 ( L) . — Morne Blue, 13.III.1921, Britton & al. 2266 ( G, K, NY) . — Heights of Aripo , 10-26. I.1922, Broadway 9929 ( K) . — Morne Blue , 18.XII.1924, Broadway 5504 ( BM, K) . — Aripo Road , 15. I.1926, Broadway 5892 ( BM, K) . — Zamsel Hill, Arouca , 15.II.1928, Broadway 6811 ( BM, K) . — Above Spring Hill, Arima-Blanchisseuse road, 26.XII.1976, Adams 14168 ( K) .

Tobago. Greenhill , 2.XII.1910, Broadway 4038 ( K, L) . — Tobago, 25.X.1925, Freeman & Williams 11426 ( K) . — Parlatuvier-Roxborough Trace, Bloody Bay to road-head above Roxborough , 8.IV.1959, Cowan 1486 ( K) . — Mount St. George to Hillsborough Dam Road , 5. VI.1975, Philcox & Raynal 7906 ( K) . — High up Roxborough to Main Ridge Road , 25.III.1993, Clement & Ryves 330 ( BM) .

Guyana. Corentyne River , IX.1879, Jenman 257 ( K) . — Near Bartica , VI.1884, Jenman 2053 ( K) . — Berbice River , XII.1886, Jenman 3593 ( K, NY) . — Pomeroon River , X.1886, Jenman 1916 ( K) . — Upper Demerara River , IX.1887, Jenman 4089 ( K, NY) . — Pomeroon River , IX.1904, Bartlett s.n. ( K, NY) . — Aruka River , X.1905, Bartlett 8576 ( K) . — Issorora, Aruka River , 16. I.1920, Hitchcock 17553 ( BM, C, K, L, NY, P, W) . — Along Potaro River, Tumatumari, VI-VII.1921, Gleason 70 ( K) . — Waini River, NW Distr., 3-18.IV.1923, De La Cruz 3663 ( K) . — Assakatta, NW Distr., 18-28. IX.1923, De La Cruz 4291 ( K) . — Marudi Mts., c. 1 km along trail from NorMan Mines camp to Aishalton , 9.XI.1982, Stoffers & al. 238 ( K) . — Region Potaro- Siparuni, Chenapou , Amerindian village ( Patumona ), 50 km upstream from Kaieteur Falls, 16.X.1987, Kvist 305 ( K) . — Potaro-Siparuni Region, trail from Kato to Paramakatoi , 12.III.1989, Hahn 5634 ( P) . — Barima- Waini Region, Aranka Head , 10.IV.1991, McDowell 4365 ( K) . — Rupununi Distr., Kuyuwini Landing and Kassikaityu River , 30.X.1992, Jansen-Jacobs & al. 3176 ( NY, P) . — Upper Ussequibo Region, Surama Village, c. 10 km N of new Parnapanum Rd. , 19.II.1990, McDowell 1889 ( FT) . — Upper Essequibo Region, Rewa River, Spider Mts. , 16.IX.1999, Jansen-Jacobs & al. 5930 ( K) .

French Guiana. La Mana, s.d., Sagot 696 ( P, W). — S.l., 1840, Leprieur. s.n. (FI-Webb, L, P). — S.l., 1842, Mélinon s.n. ( L, P) . — Cayenne-route du Mohury, 2. VI.1955, Hoock 82 ( K, P) . — Cayenne , chemin du moulin Vidal, 13. VI.1955, Hoock 89 ( K, P) . — Cayenne , 2. VI.1956, Hoock s.n. ( P) . — Près de Maripasoule , 1.IX.1961, Schnell 11692 ( P) . — Manoi , 24.VIII.1962, Hoock 93 ( P) . — Cayenne , route de Stoupan, 21. VI.1970, Oldeman B-3485 ( P) . — Tumuc Humac , frontière Brésil-Guyane, 25.VIII.1972, Granville 1380 ( P) . — Cayenne , Montravel, 8.IV.1982, Cremers 7761 ( P) . — Haute Canopi , Mont Belvédère, 23.XI.1984, Granville 6982 ( P) . — Haut Oyapock , village Zidock, 20.VIII.1985, Prévost & Grenand 1969 ( K, P) . — Île de Cayenne , Mont Baduel, 23. V.1987, Hoff 5199 ( P) . — Saül , 28.VII.1987, Hahn 3663 ( P) . — Île de Cayenne , Mont Grand Matoury, 5.IV.1995, Cremers & al. 13869 ( P) . — Bassin de l’Oyapock , Roche Touatou, 19. V.1995, Cremers & De Granville 14008 ( P) . — Montagne Trésor , le long route de Kaw, Roura, 21.XI.1997, Rouselle GU96 /106 ( MA) . — Montagne Favard , au sommet de la colline, 5.XI.1998, Poncy & Crozier 1128 ( P) . — Région de Saül , 3.IV.1997, Cremers & Crozier 14636 ( P) .

Suriname. S.loc., s.d., Moricand s.n. ( FI) . — S.loc., s.d., Hostmann s.n. ( FI) . — S.loc., s.d., Hostmann 13 ( K) . — In fruticetis Surinami, prope Lustryk , 30.XII.1837, Splitgerber 446 ( L) . — In distr. Para, II-IV.1844, Kappler 1546 ( K, W) . — Sandrij , 14-25.XI.1934, Archer 2759 ( K) . — Charlesburg Rift , 3 km N of Paramaribo, 5.IV.1944, Maguire & Stahel 24746 ( K) . — Saramacca River, trail to Coppenam River , near village, Pakka Pakka, 26. VI.1944, Maguire 23965 ( K, P) . — Inter Coppename fluv. ostium et Coronie oppidum, 15.X.1948, Lanjouw & Lindeman 1384 ( K) . — In montibus qui dicuntus Nassau , 17.II.1949, Lanjouw & Lindeman 2146 ( K) . — Jodensavanne-Mapane kreek area, along road to Mapane creek , 21.XII.1954, Lindeman 6933 ( W) . — Wilhelmina Gebergte, Zuid River , 16.IX.1963, Irwin & al. 55790a ( K, NY, P, W) . — Wilhelmina Gebergte, Kaiser Airstrip, 45 km above confluente with Lucie River , 26.IX.1963, Irwin & al. 57618, ( K, NY, P) . — Montagne Ischoli- Epoyan , 1 km du dégrad Ouarémapan, 26. VI.1972, Sastre 1460 ( P). — Nickerie, area of Kabalebo Dam Project , c. 20 km SW of Avanavero dam site, 11.XI.1976, Heyde & Lindeman 10 ( K) .

Lasiacis sloanei (Griseb.) Hitchc.

Puerto Rico. 1827, H. Wydler 222 (FI-Webb). — Toa Alta, 11.XI.1960 , R. Woodbury s.n. ( NY).

Cuba. Pinar del Rio, Mariel, 4. VI.1921, Ekman 12853 ( NY) . — Pinar del Rio, Sierra de Anafe, Loma de San Gabriel near Guanajay, 11.VII.1921, Ekman 13035 ( G) . — Oriente, Mayarì arriba, at Rio Mayarì , 7.X.1922, Ekman 15851 ( G) . — Santa Clara , Limones, Soledad , Cienfuegos, 29.VII.1927, Jack 5187 ( NY) . — Vicinity of Soledad , VIII.1941, Howard 6250 ( NY) .

Jamaica. Blue Mts. , Pimento Grove , 22. I .1913 , Harris 11589 ( BM, NY). — Orange River , near Montego Bay, 29-30.III.1920, Maxon & Killip 1673 ( NY). — Bog Walk and vicinity , 27.VII.1926, Maxon 10502 ( NY). — Middlesex , 28. I .1938 , Hunnewell 15226 ( NY). — St. Ann Parish , 1.4 miles S of Ocho Rios at end of Fern Gully, 26.XII.1971, Davidse & Convoy 32373 ( NY) .

Haiti. Port au Prince , hills between the town and Pétionville at Source Plaisance , 19.X.1924, Ekman 2190 ( C). — Vicinity of St. Louis du Nord, 30.III-7.IV 1929, Leonard & Leonard 14292 ( NY) .

Santo Domingo. Macoris , 23 km W of San Pedro de Macoris, 18.X.1946, Howard & Howard 9508 ( NY). — Jaiqui’ Picado, 20 miles W of Santiago, 27.XI.1969, Bro. Liogier 17049 ( NY) .

Antigua. Forest NE of Dark Valley, below Boggy Peak, 16.XII.1931, Box 119 ( BM, K, US: cf. Davidse 1978) . St. Martin. Pic du paradis, sommet, 424 m, 7.XII.1977, Raynal 19643 ( P) .

Suriname. Bei Paramaribo, 1913, Kruper 27915 ( L) .

Figures

FIG. 1. — C. Plumier’s unpublished manuscript of “Milium arundinaceum scandens et maculatum”: manuscript 4 in C. Plumier’s handwriting. Photo Bibliothèque centrale du Muséum national d’Histoire naturelle, Paris.

FIG. 2. — C. Plumier’s watercolour plate 82 related to manuscript 4, holotype of Panicum maculatum Aubl.(= Lasiacis maculata (Aubl.) Urb.). Photo Bibliothèque centrale du Muséum national d’Histoire naturelle, Paris.

FIG. 3. — Epitype of Panicum maculatum Aubl. (= Lasiacis maculata (Aubl.) Urb.), “Martinique, environs de St. Pierre, Mai 1853, Belanger 390” (FI-Webb).