Goeldia obscura ( Keyserling, 1878 )

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 180-181

Goeldia obscura ( Keyserling, 1878)

( Figs 18 , 25 )

Titanoeca obscura Keyserling, 1878: 591 pl. 14, fig. 15 (holotype ♀ from Santa Fé de Bogotá (4°35’56”N 74°04’51”W), Colombia, deposited in NHM, examined).

Amaurobius obscurus : Petrunkevitch 1911: 105.

Goeldia funesta : Lehtinen 1967: 236 (Part).

Remarks. The description of a male presented by Leech (1972) for Goeldia obscura refers to the specimen described by Keyserling (1891) and it was erroneously interpreted as the male of G. obscura ( Keyserling, 1878) . Here we present the first description of the male of Goeldia obscura ( Keyserling, 1878) .

Additional material examined. COLOMBIA. Huila: San Agustín (1°52’45”N 76°16’06”W), 1♂, 24.III.1994, V. & B. Roth coll. ( CAS 9025938 ); San Agustín, Archaeological site of San Agustín (1°55’N 76°14’W), 1♂, 9.IX.1980, C. Gold coll. ( CAS 9025935 ) ; Cundinamarca: Mosquera, Laguna de la Herrera (4°42’N 74°18’W), 1♀, VII.1991, E. Flórez coll. (ICN-Ar 6067); Cachipay (4°43’51”N 74°26’15”W), Alto Tolú, elev. 1600 m, 1♀, 5.XII.1996, E. Floréz coll. (ICN-Ar 1611) ; Bogotá (4°35’56”N 74°04’51”W), elev. 2600 m, 3♀, 2 juv. 20.V.1979, C. Hernandez coll. (ICN-Ar 5349); Mosquera (4°42’28”N 74°13’58”W), Mondoñedo, elev. 2600 m, 4♀, 11 juv., 6.IV.2002, C. Niño, G. Hora, J. Martinez & A. Castañeda coll. (ICN-Ar 2082); 2♀, 7 juv., 23.III.2002, J. Martinez, C. Niño, G. Mura & A. Costañeda coll. (ICN-Ar 2083) .

Diagnosis. Males of Goeldia obscura differ from other species by the short, thick, and distally constricted patellar apophysis ( Fig. 18C ); and by the tegular process distally narrowed and ventrally curved ( Fig. 18A ). Females are recognized by the wide median field of the epigynum not enclosing the copulatory openings ( Fig. 18E ); the copulatory ducts distally bent ( Fig. 18F ); the vertical spermathecae shorter than the copulatory ducts, and the horizontal spermathecae ventrally curved ( Fig. 18F ).

Description. Male (CAS 9025935): Coloration: carapace orange, palp brown and femora black, other leg segments yellowish brown. Total length 4.9. Carapace: length 2.25, width 1.65, height 0.68. Opisthosoma: length 2.25, width 2.2. Clypeus: height 0.16. Eye diameters and interdistances: AME 0.05; ALE 0.10; PME 0.08; PLE 0.06; AME–AME 0.11; AME–ALE 0.11; ALE–PLE 0.05; PME–PLE 0.2; PME–PME 0.13; AME–PME 0.13. Chelicerae paturon: length 0.10. Leg measurements: I: Femur 1.73/ patella 0.65/ tibia 1.68/ metatarsus 1.63/ tarsus 0.88/ total 6.55; II: 1.75/ 0.65/ 1.63/ 1.45/ 0.68/ 6.15; III: 1.50/ 0.63/ 1.38/ 1.23/ 0.68/ 5.40; IV: 1.70/ 0.68/ 1.80/ 1.45/ 0.63/ 6.25; Leg formula: 1423; Leg spination: femur I–II: p1ap; tibia II v0-0-1, p1ap; III v1-0-1 or v1-0-2, p1ap, r1ap; IV v1-0-1 or v0-1-2, p1ap, r1ap; metatarsus I v2-2- 1m, p1ap, r1ap; II v2-2-1map, r1ap, p2ap; III v2-2- 1m, p1-1-1ap, r0-0-1ap, d1-0-1rap or d0-0-1p; IV v2-2-2- 1m, p0-2-2ap, r1ap, d2ap. Patellar apophysis dorsally projected ( Fig. 18B–D ). MLT with apex membranous ( Fig. 18C ). Spermatic duct bending at 12 o’clock. Pars pendula thickening at the tip of the embolus ( Fig. 18D ).

Female (CAS 9025998). Coloration: carapace brown with the edge darker; chelicerae, labium sternum and endites brown; coxae and femora brown; other leg segments yellowish brown; opisthosoma pale brown. Total length: 5.40. Carapace: length 2.1, width 1.6, height 1.03. Opisthosoma: length 3.5, width 2.2. Clypeus: height 0.18. Eye diameters and interdistances: AME 0.07; ALE 0.11; PME 0.10; PLE 0.10; AME–AME 0.10; AME–ALE 0.13; ALE–PLE 0.06; PME–PLE 0.23; PME–PME 0.15; AME–PME 0.11. Chelicerae paturon: length 1.06. Leg measurements: I: femur 1.56/ patella 0.59/ tibia 1.53/ metatarsus 1.22/ tarsus 0.72/ total 5.63; II: 1.47/ 0.69/ 1.34/ 1.09/ 0.69/ 5.28; III: 1.34/ 0.66/ 0.97/ 0.94/ 0.50/ 4.41; IV: 1.69/ 0.75/ 1.47/ 1.09/ 0.53/ 5.53. Leg formula: 1423. Leg spination: femur I p1ap; metatarsus I–III v1map, p1ap, r1ap; IV v0-1-1map, r1ap, p1ap. Median field of epigynum wider than long but not covering the copulatory openings ( Fig. 18E ). Fertilization ducts apart from each other twice its length ( Fig. 18E–F ).

Variation. Length (10 ♀): total 4.82–6.15; carapace 1.8–2.6; femur I 1.48–1.70.

Distribution. Colombia ( Fig. 25 ).

Figures

FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities.