Goeldia diva, Almeida-Silva & Brescovit, 2024

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 163-165

Goeldia diva sp. nov.

( Figs 9 ; 25 )

Type material. Holotype ♂ from Olivença (14°56’48”S 39°0’43”W), Ilhéus, Bahia, Brazil, 25/VI/1998, no coll. ( IBSP 35919 ) . Paratypes: 1♀ from Fazenda São Roque , Gandu (13°44’38”S 39°29’13”W), Bahia, Brazil, Equipe CEPLAC coll. ( MNRJ 3102 ) ; 2♀, 2♂ from Reserva Michelin, Ituberá (13°43’55”S 39°08’56”W), Bahia, C. M. Pinto-Leite coll. ( MZUFBA 2440–2443 ) .

Additional material examined. BRAZIL. Bahia: Salvador , Bahia de Aratu (12°47’32.8”S 38°28’15.3”W), 3♀, 2007–2008, K. Benati coll. ( IBSP 123565–123567 ) ; Itamaraju (17°02’20”S 39°31’51”W), Fazenda Santa Úrsula, 1♂, 2♀, no collector or date ( MNRJ 3189 ; 13392); Gandu (13°44”38”S 39°29’13”W), Fazenda São Roque , 1♂, 1♀, 21.XI.1969, CEPLAC team coll. ( MNRJ 1916 ; 3201); Gandu (13°44”38”S 39°29’13”W), 1♂, 1978-1979, no collector ( MNRJ) ; Same, Fazenda Pedra Branca, 2♂, 3♀, 22.X.1979, CEPLAC team coll. ( MNRJ 2618 ; 3201; 2733; 3042); Camacan (15°25’08”S 39°29’45”W), 3♂, 2♀, CEPLAC team col. ( MNRJ 3174 ; 2886); Same, Fazenda Matiapã , 1♂, 14.X.1978, J.S. Santos coll. ( MCN 20254 ) ; Jussari (15°11’27”S 39°29’42”W), 2♂, 2♀, CEPLAC team coll. ( MNRJ 3143 ; 3296; 13316; 3131) .

Etymology. The specific epithet is an honor to LMAS’s sister, Elisabete Almeida (Bete Diva), who is a professional opera singer and performer. The word entered the English language in the late 19th century and it is derived from the Italian noun diva , a female deity or goddess. The meaning of diva is closely related to prima donna, Italian for “first lady”, the leading female singer in an opera company.

Diagnosis. Goeldia diva sp. nov. differs from the other species by the tegular process long, with an acute tip and broad membranous base ( Fig. 9A ); and by the patellar apophysis thick and posteriorly curved ( Fig. 9B–D ) on the male palp. The female can be distinguished by the epigynum wide but with lateral rims not covering the copulatory openings ( Fig. 9E ); horizontal spermathecae distally projected anteriorly ( Fig. 9F ); vertical spermathecae with the same height as the copulatory ducts and distally curved ( Fig. 9F ).

Description. Male (IBSP, holotype). Coloration: carapace, chelicerae, labium, endites and femora dark brown; coxa and other leg segments yellowish-brown; opisthosoma grayish brown. Total length 4.60. Carapace: length 2.05, width 1.65, height 0.80. Opisthosoma: length 2.20, width 1.55. Clypeus: height 0.12. Eye diameters and interdistances: AME 0.09; ALE 0.01; PME 0.01; PLE 0.09; AME–AME 0.01; AME–ALE 0.09; PME–PME 0.09; PME–PLE 0.12; AME–PME 0.10; ALE–PLE 0.04. Paturon: 1.09 long. Leg measurements: I: femur 2.21/ patella 0.65/ tibia 2.00/ metatarsus 1.90/ tarsus 0.92/ total 7.70; II: 2.04/ 0.82/ 1.84/ 1.63/ 0.76/ 7.08; III: 1.8/ 0.73/ 1.5/ 0.67/ 0.64/ 6.36; IV: 2.09/ 0.66/ 1.93/ 1.75/ 0.68/ 7.13. Leg formula: 1423. Leg spination: femur I–II: p 1ap; tibia II v1r-0-1-p1ap; III v1-0-2, p1ap, r1ap; IV r1ap, v0-0-1; metatarsus I v2-1p-1map, p1ap, r1ap; II v2-1p-1map, p2ap, r1ap; III v1-2-1 or v2-2-1, p1-1-1, r0-1-1, d1msa; IV v1ap, p1ap, r1ap or r2ap; v2-2-1, p2ap, r1ap; tarsus IV v0-1-0. MLT apex totally membranous ( Fig. 9C ). Tegular process shaped as Fig. 9A . Spermatic duct terminal fold pointing upward ( Fig. 9B ). Pars pendula thickening far from the embolic tip ( Fig. 9D ).

Female (MNRJ 3102, paratype). Coloration: carapace with cephalic area slightly darker than thoracic area; chelicerae reddish brown; labium, endites and leg femora brown; other leg segments reddish brown except in leg IV, in which the apex of the segments is slightly lighter. Total length: 5.50. Carapace: length 2.60, width 1.60, height 0.78. Opisthosoma: length 2.95, width 2.0. Clypeus: height 0.18. Eye diameters and interdistances: AME 0.06; ALE 0.14; PME 0.11; PLE 0.14; AME–AME 0.09; AME–ALE 0.13; PME–PME 0.14; PME–PLE 0.21; AME–PME 0.12; ALE–PLE 0.06. Paturon: 1.06 long. Leg measurements I: femur 1.74/ patella 0.76/ tibia 1.62/ metatarsus 1.35/ tarsus 0.83/ total 6.30; II: 1.56/ 0.75/ 1.31/ 1.16/ 0.7/ 5.49; III: 1.48/ 0.72/ 0.98/ 1.04/ 0.52/ 4.74; IV: 1.78/ 0.78/ 1.50/ 1.24/ 0.51/ 5.81. Leg formula: 1423. Leg spination: metatarsus: I to III: p 1ap, r 1ap, v 1ap; IV: v 0-1-1, r 1ap, p 1ap. Posterior edge of the epigynum median field sinuous ( Fig. 9E ). Small portion of the copulatory duct wall visible near the copulatory opening ( Fig. 9E ). Epigynal rim very small ( Fig. 9E ). Copulatory ducts very sclerotized ( Fig. 9F ) and partially visible by transparency ( Fig. 9E ).

Variation. Length (10 ♂): total 3.90–4.80; carapace 2.00–2.30; femur I 1.80–2.30; (10 ♀): total 4.90–6.30; carapace 1.90–3.10; femur I 1.20–2.00.

Distribution. Northeastern Brazil: only known from the coastline of the state of Bahia, south to Salvador ( Fig. 25 ).

Figures

FIGURE 9. A–F. Goeldia diva sp. nov. A–D: male, left palp. E–F: female epigynum. A. Tegular process, frontal view. B. Ventral view C. patella and tibia, dorsal view; D. Retrolateral view; E-F. Female epigynum: E. Ventral view; F. Dorsal view.

FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities.