Upeneus randalli Uiblein & Heemstra, 2011

Uiblein, Franz, Gledhill, Daniel C., Pavlov, Dimitri A., Hoang, Tuan Anh & Shaheen, Shaker, 2019, Three new goatfishes of the genus Upeneus (Mullidae) from the Indo-Pacific, with a redescription of colour patterns in U. margarethae, Zootaxa 4683 (2), pp. 151-196 : 188-190

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Upeneus randalli Uiblein & Heemstra, 2011


Upeneus randalli Uiblein & Heemstra, 2011

Randall’s goatfish

( Figures 1 View FIGURE 1 , 2 View FIGURE 2 , 5 View FIGURE 5 ; Tables 2, 5, 6, 9)

Upeneus randalli Uiblein & Heemstra, 2011 . Type locality off S Kuwait, Arabian/Persian Gulf, Kuwait.

Holotype. Arabian / Persian Gulf : BPBM 33180 View Materials , HT, adult, 101 mm SL, Arabian / Persian Gulf, Kuwait, off S Kuwait, 29˚00’ N, 48˚25’ E, RV Bahith, trawl, 15–20 m depth (fresh-colour photo)

Paratypes (7 adults: 66–106 mm SL; 1 fresh colour photo, 1 fresh-colour drawing): Arabian / Persian Gulf: BPBM 21201 View Materials , 6 View Materials : 66-88 mm SL, Bahrain, fish market (fresh-colour photo of 80 mm SL fish); Gulf of Oman: ZMUC P 49161 View Materials , 106 mm SL, S Iran, Chahabar (fresh-colour drawing in Blegvad & Løppenthin (1944))

Non-type: Arabian/Persian Gulf: BPBM 29498, subadult, 60 mm SL, Bahrain (fresh-colour photo)

Diagnosis. Dorsal fins VIII + 9, the first spine minute; pectoral fins 13–14; gill rakers 6–7 + 17–19 = 23–25; lateral-line scales 28–30; measurements in % SL, adults: body depth at first dorsal-fin origin 23–24; body depth at anus 19–22; caudal-peduncle depth 9.9–11; caudal-peduncle width 2.9–3.8; maximum head depth 19–21; head depth through eye 15–16; suborbital depth 8.0–10; interorbital length 7.2–8.8; head length 27–30; snout length 11–12; postorbital length 10–12; orbit length 6.2–7.4; upper-jaw length 10–11; barbel length 16–20; interdorsal distance 13–15; caudal-peduncle length 23–25; caudal-fin length 27–30; anal-fin height 15–19; pelvic-fin length 20–22; pectoral-fin length 20–22; pectoral-fin width 3.8–4.4; first dorsal-fin height 19–22; second dorsal-fin height 16–20; measurements in % SL, subadults (based on single specimen): body depth at first dorsal-fin origin 22; body depth at anus 19; caudal-peduncle depth 9.0; caudal-peduncle width 3.5; maximum head depth 18; head depth through eye 16; suborbital depth 7.0; interorbital length 7.7; head length 28; snout length 12; postorbital length 10; orbit length 8.2; upper-jaw length 11; barbel length 19; interdorsal distance 14; caudal-peduncle length 23; caudalfin length 29; anal-fin height 16; pelvic-fin length 21; pectoral-fin length 20; pectoral-fin width 3.7; first dorsal-fin height 21; second dorsal-fin height 19; fresh colour: head and body dorsally red-brown red or grey mottled, ventrally white or rose, in some fish with tiny red or brown dots; caudal fin with 11–13 (9 in subadult) brown or darkgrey oblique bars crossing both lobes; on upper caudal-fin lobe 4 or 5 bars, on lower lobe 5–8 bars; lower caudal lobe may be longitudinally crossed by a dark grey stripe (only observed in single subadult); caudal-fin bars and hyaline interspaces of about pupil width in adults; in subadult hyaline interspaces on upper lobe and bars on lower lobe wider (based on single subadults); lower caudal-fin lobe tip sometimes black; barbels white; a single yellow, pale beige or pale red mid-lateral body stripe of less than pupil width from behind head to caudal peduncle, the stripe covered by 3 sections with horizontal series of 2–4 dark brown or black dots, the posteriormost series of 2 dots behind second dorsal-fin base connecting to a dark saddle placed behind second dorsal fin; first dorsal fin with closely fused pale-red brown or pale-grey stripes; the tip region of first dorsal fin without dark pigmentation; second dorsal fin with 2–3 well-separated, brown or black stripes with hyaline interspaces, the dorsal-most stripe the largest and darkest; preserved fish brown or pale brown, mid-lateral body stripe completely lost, often with remains of dark saddle and—more rarely—other dark pigmentation patterns.

Distribution, habitat and size. Arabian/Persian Gulf and inner Gulf of Oman: depth 15–20 m; maximum size 11 cm SL.

Intraspecific comparison. The single subadult differs from adults in shallower maximum body and head depth, shallower caudal peduncle, shallower suborbital depth, larger eyes, longer anal-fin base, slightly narrower pectoral fins and wider bars on caudal fin, especially on lower fin lobe which is crossed by a dark stripe at mid of lobe ( Tables 2, 9; Figure 2 G, H View FIGURE 2 ).


Collection of Leptospira Strains


Zoological Museum, University of Copenhagen


Bishop Museum














Upeneus randalli Uiblein & Heemstra, 2011

Uiblein, Franz, Gledhill, Daniel C., Pavlov, Dimitri A., Hoang, Tuan Anh & Shaheen, Shaker 2019

Upeneus randalli

Uiblein & Heemstra 2011