Phytocrene borneensis Becc.

Rio, Cédric Del, Stull, Gregory W. & Franceschi, Dario De, 2020, Survey of the fruits and endocarps of Icacinaceae (Lamiids, Icacinales), European Journal of Taxonomy 645, pp. 1-130 : 62

Phytocrene borneensis Becc.

Fig. 21.1–21.8

Material examined

Specimen used for endocarp and fruit description

MALAYSIA • Borneo , Sarawak; 8 Jul. 1979; B. Lee S.40246; L [ L.2289394] .

Description

FRUIT. Obovoid, slightly asymmetrical and widely acuminate at the apex, yellow-brown when mature. Epicarp with clusters of yellow hairs. Mesocarp thin but thick at the accrescent part when dry. Calyx not persistent. Length ca 56 mm, width ca 14 mm.

ENDOCARP. Brown, obovoid in lateral view, lenticular in transverse section, length ca 39.1 mm, width ca 17.3 mm, thickness ca 9 mm. No clear keel surrounding the endocarp in the plane of symmetry, but thin trace of keel present in the upper part. Apex asymmetrical and emarginate in lateral view; base acute, symmetrical. Outer surface of the endocarp pitted. Pits circular in the upper half and more elongate in the lower half of the endocarp, relatively thin, 0.2–0.4 mm in width, randomly arranged with ca 12–13 pits longitudinally and 10–11 pits transversally (ca 125–130 pits per face). Pits narrow, without tubercles. Endocarp wall 681–766 µm thick excluding pits (307–351 µm thick including pits). Endocarp wall (excluding pits), with three cell layers: outermost layer with ca 16 rows of anticlinally oriented cells, cells 32.8–45.7 µm in length, followed by a layer with ca 9 rows of periclinally oriented cells, cells 14.5–24.0 µm in width; innermost layer consist of periclinally oriented cells, cells 9.7–11.4 µm in width, lining the locule surface with inflated cells. Locule covered with rounded protuberances due to the thickening of the wall under the pits.