Makromastax, Santos-Silva & Botero & Tirant, 2018

Santos-Silva, Antonio, Botero, Juan Pablo & Tirant, Stéphane Le, 2018, A new genus, four new species and taxonomic and geographic notes in Mexican Cerambycidae, Zootaxa 4420 (2), pp. 189-210: 201

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gen. nov.

Makromastax   , gen. nov.

Type species: Haplidus mandibularis Chemsak & Linsley, 1963   , monotypy, present designation.

Etymology. A compound of the Greek µᾰκρός (makrós, “long”) + ΜΑστάξ (mástax, “jaws”); refers to the long and falciform mandibles in males, the more conspicuous characteristic of the new genus. Feminine gender.

Description. Body flattened; head prognathous; frons moderately short, transverse, nearly vertical; antennal tubercles slightly elevate with rounded apex; palpi slender, maxillary palpi long, almost reaching apex of mandible (when closed) in male, surpassing in female; labial palpi distinctly shorter than maxillary palpi; postclypeus large, nearly perpendicular to frons, flap-shaped, entirely hiding labrum in dorsal view. Mandibles in male falciform, distinctly longer than half of head, apex bifid, superior margin of inner side carinate, ending abruptly near apex, inner surface excavated, inferior inner margin with large tooth before middle, outer side without tooth; in female with similar general appearance, but distinctly shorter. Upper eye lobes with 5 rows of ommatidia at its maximum width; distance between them equal to about 4 times width of a lobe in male, slightly shorter in female. Antennae 11-segmented, slightly longer than body in male, about as long as body in female; scape elongate, slightly, gradually widened toward apex; antennomeres without apical spicule; antennomere III not carinate dorsally, longer than IV, shorter than scape; basal antennomeres with long, erect, sparse setae ventrally. Prothorax distinctly wider than long; sides broadly rounded, unarmed. Pronotum without gibbosities. Prosternal process gradually narrowed toward apex. Procoxal cavities widely open laterally. Mesoventral process slightly narrowed toward apex; apex without lateral lobes. Elytra parallel-sided, not carinate; apex rounded, unarmed. Femora fusiform, slightly pedunculate basally in meso- and metafemora. Tibia not carinate laterally. Metatarsomere I about 1.5 times II –III together. Abdominal ventrites with pubescence not obscuring integument.

Remarks. Makromastax   gen. nov. differs from all other American Hesperophanini   by the long and falciform mandibles in male. Chemsak and Linsley (1963) provided a key to Mexican genera of Hesperophanini   , but including genera currently placed in other tribes ( Eburia Lacordaire, 1830   , and Eburodacrys White, 1853   , Eburiini   ; Chlorida Audinet-Serville, 1834   , Bothriospilini   ; Amphelictus Bates, 1884   , Cerambycini   ). Two other genera were described after Chemsak and Linsley (1963): Megosmidus Hovore, 1988   , and Oraphanes Chemsak and Linsley, 1984   .