Siphona (Aphantorhaphopsis) coactilis, Tachi & Shima, 2023

Tachi, Takuji & Shima, Hiroshi, 2023, Systematic study of the subgenus Aphantorhaphopsis of the genus Siphona in East Asia and the Oriental Region (Diptera: Tachinidae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (1), pp. 213-247 : 220

Siphona (Aphantorhaphopsis) coactilis sp. nov.

( Figs 11–15 , 88 )

Type material. HOLOTYPE: J ( SMNH), MALAYSIA, Pahang / Bukit Fraser , 1200 m / Malaise-trap, in jungle / 2– 6.04.1992./ H & H. Hippa . PARATYPES: MALAYSIA: PENINSULA: PAHANG: 30 JJ, same data as holotype; 13 JJ 8 ♀♀, Cameron Highlands, Gunung Jasar, 1700 m, 20–23, 24–27.xi.1994, T. Pape. BORNEO: SABAH: 1 J 1 ♀, Kinabalu Park, 1600 m, 2–9.xi.1994, T. Pape (all in SMNH).

Diagnosis. Fronto-orbital plate with brownish pruinosity and with a row of short setae on lower part; frontal vitta reddish brown; 3 postsutural dorsocentral setae; abdomen with whitish pruinosity.

This species is similar to S. (A.) kanmiyai sp. nov. in general appearance but differs from it in a row of short setae on the fronto-orbital plate.

Description. Body length 3.5–4.0 mm. Male. Head whitish in ground color; fronto-orbital plate with brownish pruinosity; frontal vitta reddish brown; antenna with scape and pedicel light brown; postpedicel dark brown; basal half of arista light brown, blackish in apical half; palpus yellow. Vertex approximately 0.34 of head width; parafacial narrower than width of 2nd aristomere at middle height; gena approximately 0.22 of eye height; anterior reclinate orbital seta posterior to middle of fronto-orbital plate; fronto-orbital plate with row of short setae on lower part; antenna with postpedicel subrectangular, approximately twice as long as wide, and approximately 4 times as long as pedicel; 2nd aristomere approximately 3 times as long as wide; 3rd aristomere thickened in basal 1/3; palpus clavate; prementum approximately 4 times as long as wide and approximately half as long as eye height; labella pad-like.

Thorax. Dorsum slightly dark gray, with yellowish pruinosity; apical 1/3 of scutellum orange; pleura brown, with gray pruinosity. Three presutural and 3 postsutural dorsocentral setae.

Wing. Tegula black; basicosta orange. Relative lengths of costal sectors two, three, and four approximately 1: 7: 3; ultimate section of M 4 approximately 0.3 times as long as penultimate section, and subequal in length to crossvein dm-m; R

1

bare.

Legs yellow in ground color; tarsi blackish. Fore tibia with 2–4 anterodorsal, 3–4 posterodorsal and 1 posterior setae; hind tibia with 2–4 anterodorsal, 2–4 posterodorsal and 3–5 ventral setae.

Abdomen yellow in ground color, with whitish pruinosity on anterior margin of tergites 3–5; posterior 1/2–4/5 of tergites 3–5 blackish. Syntergite 1+2 without lateral marginal setae; tergite 3 with pair of lateral and median marginal setae; sternite 5 with pair of relatively rounded median lobes on inner edge.

Male postabdomen. Surstylus somewhat narrowed in middle in lateral view and longer than cerci; cerci nearly entirely thickened in lateral view; postgonite curved ventrally in middle and rounded apically; pregonite somewhat pointed apically in lateral view, without seta; epiphallus absent; distiphallus mostly sclerotized in lateral view, with some tiny spinules ventrally.

Female. Differing from male as follows: gena approximately 0.17 of eye height; postpedicel approximately 3.5 times as long as wide.

Female postabdomen. Tergites 6–7 absent; sternite 7 nearly fan-shaped with a short anterior apodeme.

Etymology. The name “ coactilis (made thick)” is taken from the cerci of the male postabdomen; an adjective.

Host. Unknown.

Distribution. Malaysia (Pahang, Sabah).

Figures

Figs 11–15. Male and female postabdomen of Siphona (Aphantorhaphopsis) coactilis sp. nov. (paratype, Taiwan, Taichung). 11 – epandrium, cerci and surstylus in lateral view; 12 – epandrium, cerci and surstylus in dorsal view (setae omitted on left side); 13 – hypandrium, pregonite, postgonite and phallus in lateral view; 14 – male fifth abdominal sternite in ventral view (setae omitted on left side); 15 – female postabdomen in lateral view. Scale bars = 0.1 mm.Abbreviations: cer – cerci; ep – epandrium; hy – hypandrium; hyp – hypoproct; ph – phallus; pos – postgonite; pre – pregonite; st – abdominal sternite; sur – surstylus; t – abdominal tergite.

Figs 86–91.Adult male habitus of Siphona (Aphantorhaphopsis) spp. 86 – Siphona (Aphantorhaphopsis) expleta sp. nov. (paratype, Philippines); 87 – S.