Pseudoneureclipsis Ulmer

Oláh, János & Johanson, Kjell Arne, 2010, Contributions to the systematics of the genera Dipseudopsis, Hyalopsyche and Pseudoneureclipsis (Trichoptera: Dipseudopsidae), with descriptions of 19 new species from the Oriental Region., Zootaxa 2658, pp. 1-37 : 21-22

Pseudoneureclipsis Ulmer

Pseudoneureclipsis Ulmer, 1913: 84 . Type species: Pseudoneureclipsis ramosa Ulmer (monotypic); type locality: Indonesia (Java).

Ulmer (1913) originally placed this genus in the Polycentropodidae because it had the combination of wing venation characters of other then-known polycentropodids: (1) forewing lacking fork V; (2) forewing lacking an additional costal crossvein; and (3) hind wing with fork III, the latter character relating it to Neureclipsis . All species described below exhibit the following characters:

Adult. Head. In dorsal view, head almost quadrangular and eyes about half as wide as vertex width. Maxillary palp formula II-I-III-IV-V. Antennae stout, 1.4 times as long as forewings; scapes short, about 1/4th as long as head; distance between antennae longer than length of scapes. Labrum bipartite, posterior part sclerotized, anterior part membranous. Mandibles enlarged, broad, surfaces granulated papillose. Some species with modified mandibles as described below. Laciniae elongate setiferous, curving mesad. Anterior tentorial pits located laterally near eyes. Pregenae narrow. Clypeus and frons wide; epistomal suture connecting tentorial pits nearly invisible; theoretical line separating clypeus and frons detectable by setose frons and bare clypeus. Frontal grooves almost indiscernible; possibly represented by heavily sclerotized posterior margin of bulging frons near antennal sutures of antennal sockets. Vertical section of sclerotized lateral margin of frontoclypeus constituting frontogenal grooves above anterior tentorial pits and clypeogenal grooves below anterior tentorial pits. Clypeolabral groove well-developed, encircling mandibles. Coronal groove (stem of epicranial groove) forming mid-line along vertex and occiput. Occipital grooves well developed, forming unique primary intersegmental suture; postoccipital groove visible between occipital warts. Pairs of vertexal lateroantennal setose warts and vertexal ocellar compact setose warts differentiated. Vertexal medioantennal and vertexal medioocellar diffuse setose warts partially fused into mesal bands on vertex. Pair of occipital compact setose warts dominating on posterodorsal region of head. Narrow postgenal compact setose warts present, visible in caudal and ventral views. Pair of vertexal tubercles, sensory papillae or sensillae basiconica present posterior of crossing of coronal and occipital grooves. Cervical sclerites forming narrow, long, rod-like anterior arms articulating anteriorly with elongate occipital condyles immediately above posterior tentorial pits. Cervical sclerites fused with posterior cervical sclerites. Posterior cervical sclerites about half as long as anterior cervical sclerites; forming triangular plates; each with one corner reaching and articulating with prothoracic episternum; other 2 corners forming rod-like ventral intercervical sclerites articulating to weakly sclerotized anteromedian band of prothoracic eusternum. Cervical sclerites without setae. Large cervical setose warts attached to dorsum of anterior, rod-like, sclerites near posterior cervical sclerites.

Thorax. Two pairs of large, rounded, pronotal compact setose warts; lateral pronotal warts slightly smaller than mesal warts; mesal warts nearly tangential mesally, separated by narrow gap. Pair of mesoscutal compact setose warts rounded, indistinct; pair of longitudinally elongate mesoscutellar compact setose warts slightly triangular, narrowing posterad, nearly occupying mesoscutellar area. Spur formula 2, 4, 4. Each foreleg posterior spur being half as long as anterior spur; mid and hind leg posterior spurs about 1/3rd as long as anterior spurs. Leg claws small, symmetrical. Forewing membrane and veins light brown, covered by scarce cinereus setae. Forewing forks I, II, III and V present; discoidal cell, median cell and thyridial cells closed; crossvein m-cu not reaching point of bifurcation of M; and forewing fork I smaller than fork III, except when otherwise stated. Hind wing forks II, III and V present; discoidal and median cells open; thyridial cell closed, narrow.

Remarks. The detailed head and thoracic structures, including tentorium, grooves, sutures and setal warts of the genus are described for the first time below in P. quancong , new species, and proved stable in the genus. However, the sclerites, spines and microtrichia of the phallic apparatus were difficult to describe precisely. There is a wide size range of the phallic microtrichia, from almost invisible to large spine-like structures. In addition, there are probably indistinctly discernible phallotremal sclerites among the pale structures. A standard description of the microtrichia structure is hindered by the erection and eversion state of the endotheca. All phallic processes (rod-like processes of tergite and sternite IX, preanal appendages, and paraproctal processes) meet at the lateral fulcra. The phallic apparatus is fixed to this complex by paraproctal straps.