Otiorhynchus, Germar, 1822, Germar

Christoph Germann, 2016, A new Otiorhynchus Germar, 1822 subgenus Tecutinus Reitter, 1912 from Anatolia (Coleoptera, Curculionidae), Contributions to Natural History 33, pp. 1-10 : 5-9

Key to species (males are required; the characters of O. crassicornis GYLLENHAL, 1839 are based on Stierlin's redescription (1861)). For distribution see Fig. 3 .

1. Tip of penis pointed and elongated tongue-like .......................................... 2

– tip of penis pointed, cut or rounded ............................................................ 3

2. Disc of pronotum densely punctured, pronotum transverse; interspaces on elytra leather-like, finely punctured, striae shallow. Rostrum short, rectangular, pterygia narrow; antennae thicker; metatibiae in males less deeply incised, margin not carinate (6.8–7.5 mm). Lebanon [Pass between Ainata and Becharré; Mount Sannin above Beyrouth].............................................. ..................................................................... kindermanni STIERLIN, 1861

– disc of pronotum tuberculate, tubercles flat, irregular and intermixed with punctures; rostrum longer than wide; elytra chagrinated; antennae slenderer; metatibiae of males more deeply incised on inner side before apex, margin granulate (7.0–9.5 mm). Monte Salbakos [=Karci Dağları]................. ................................................................................... catonii LoNA, 1943

3. Tip of penis pointed..................................................................................... 4

– tip of penis cut or rounded .......................................................................... 5

4. Habitus robust with broad elytra, strongly tuberculate pronotum and simply pointed tip of penis (8.5–10.3 mm). Crete Island ..................................... ........................................... lefkaoriensis GERMANN & CoLoNNELLI, 2015

– habitus gracile, elytra elongate oval, disc of pronotum punctured, tip of penis pointed and bispinate (7.7–8.5 mm). Ak Dağları .................................. .............................................................................. charleshuberi sp. nov .

5. Elytra with fine grey hairs............................................................................ 6

– elytra without fine grey hairs .................................................................... 10

6. Small species (6.2–6.5 mm); vestiture of elytra consisting of evenly distributed dense short grey hairs. Giresuni Dağları ............................................... ........................................................................ torulensis Benedikt, 2000

– bigger species (>8.5 mm); vestiture not homogenous but patchy or intermixed with dark bristles .............................................................................. 7

7. Vestiture of elytra with patches of grey hairs without dark bristles; inner sides of protibiae in males scabrous and with tooth in the middle (9–10 mm). Eskişehir........................................................ escherichi REITTER, 1898

– vestiture with grey hairs and dark bristles; inner sides of protibiae in males without tooth in the middle ......................................................................... 8

8. Pronotum transverse with shiny tubercles, laterally strongly rounded; rostrum longer, rostral dorsum strongly furrowed (size not known). "Persia occidentalis" [most likely Dagestan]............... crassicornis GYLLENHAL, 1839

– pronotum less transverse with flattened dull tubercles, laterally less rounded; rostrum shorter, rostral dorsum flat ............................................ 9

9. Rostral dorsum short and well separated from head, rostrum forming an equilateral triangle; metatibiae in males less deeply incised on inner margin before apex (8.5–9 mm). Bulgar Maaden [= Bolkar Dağları].................... ....................................................... crinitellus K. Daniel & Į. Daniel, 1902

- head less separated from rostrum, rostrum longer, forming an acute triangle; metatibiae in males more deeply incised on inner margin before apex (9–10.5 mm). Iskilib.......................................... paracrinitellus BRAuN, 1989

10. Striae on elytra with coarse grooves ......................................................... 11

– striae not groove-like deepened ............................................................... 13

11. Grooves very broad and irregular, overlapping in first two rows and in rows 3 and 4, therefore first and third interspaces not visible; disc of pronotum densely set with flat tubercles, not punctured; metatibiae of males strongly incised on inner margin before apex; apex of fore tibiae clearly protruding outwards in both sexes (8.3–10.3 mm). Mihaliccik .................................. ........................................................................... heinzianus Braun, 1988

– grooves on elytra small, standing in regular rows, first and third interspaces visible; disc of pronotum punctured; apex of fore tibiae straight or weakly protruding outwards ..................................................................... 12

12.Pronotum narrow; meso- and metatibiae in males only weakly broadened, metatibiae inconspicuously incised (7.5–9.5 mm). "Gebirge Armeniens" [from NE Turkey (Artvin) to central Caucasus, Armenia] ................................ ................................................................... cribripennis HocHHuTH, 1851

– pronotum broader; meso- and metatibiae in males strongly broadened towards apex, inner sides of metatibiae strongly incised before apex (8.5– 10 mm). "Amasia" ..................................... pseudocribripennis BRAuN, 1988

13.Elytral striae strong, pronounced.............................................................. 14

– elytral striae shallow or hardly visible ....................................................... 17

14.Disc of pronotum and sides mainly with flattened tubercles, only scarce and isolated punctures in between ........................................................... 15

– disc of pronotum mainly punctate, small tubercles towards sides of pronotum ............................................................................................................ 16

15. Antennae short and thick with transverse funicular segments; smaller species (<8 mm); pronotum broad, laterally strongly rounded; tip of penis somewhat narrowed towards tip, with straight fore margin (7.8–8 mm). Ikisdere .................................................... ikisderensis SMRECzYÑsKI, 1970

– antennae slenderer with funicular segments less transverse; bigger species (>9 mm); pronotum narrower, laterally less rounded; penis rounded (not cut) at tip (9–10.5 mm). Sivas Kizildag Gecidi ........................................ ................................................................................. riedeli BRAuN, 1989

16.Antennae slenderer, funicular segments less transverse; metatibiae in males less deeply incised on inner margin before apex; penis more rounded at tip (8.2–8.5 mm). Cesarea [Erciyas Dagi] .............................................. ....................................................................... caesareus K. DANIEL, 1906

– antennae thicker, funicular segments more transverse; metatibiae in males more deeply incised on inner side before apex; penis cut, almost angular and faintly divergent just before tip (7.1–8 mm). Soganli Dagi ...................... ............................................................. soganliensis SMRECzYÑsKI, 1977

17. Surface of elytra without chagrination, leather-like with tiny sharp tubercles on interspaces towards sides and declivity........................................ 18

– elytral surface chagrinated (inconspicuous only in karagolensis ), with tiny tubercles or punctures on interspaces ...................................................... 21

18. Frons deepened; body shorter; pronotum more gracile, much smaller than elytra; protibiae almost straight, only weakly granulated; metatibiae in males deeply incised on inner side before apex (7–9.9 mm). Ilgaz Gecidi ..... .............................................................................. fremuthi BRAuN, 1989

– frons vaulted; body massive, oval to broad oval; pronotum stronger; metatibiae in males either deeply incised on inner side before apex (tmolosensis) or only moderately incised ......................................................... 19

19. Eyes strongly protruding; apex of protibiae only weakly protruding outwards; metatibiae of males on inner margin deeply incised before apex; elytra at disc flattened, declivity less steep (10.5–11 mm). Tmolos Dağları [=Boz Dag]............................................................... tmolosensis LoNA, 1943

– eyes less protruding; apex of protibiae strongly protruding outwards ( Arammichnus -like); elytra at disc convex; declivity steeper .................... 20

20.Eyes smaller; protibiae emarginate on inner side with several tooth-like tubercles along the middle, apex protruding outwards (8–9.5 mm). Sultan Dağlar........................................... gymnopterus K. DANIEL & J. DANIEL, 1902

– eyes bigger; protibiae without tooth-like tubercles on inner side, only weakly granulate (8–9.6 mm). Tarsus [= Mersin]; Camliyayla........................ ........................................................................... latifrons STIERLIN, 1890

21. Bigger species (>10 mm) with massive body; metatibiae of males broadened but not cut on inner side before apex (10 –10.5 mm). Monte Salbakos [= Karci Dağları] .................................................... salbakosanus LoNA, 1943

– smaller species (<8.5 mm); body less massive.......................................... 22

22. Antennae short and thick; funicular segments transverse; microsculpture of elytra less apparent; metatibiae with long spur-like apex on inner side (6.7–8.5 mm). Giresun Dağları.................. karagolensis SMRECzYÑsKI, 1970

– Antennae slenderer, only some funicular segments transverse; microsculpture of elytra more apparent; metatibiae without spur ............................. 23

23. Antennae with funicular segments from fourth on transverse; elytra oval; apex of metatibiae in males moderately incised on inner side before apex (6.2–7.8 mm). "Constantinopolis" [Bithynian Olymp; Uludag]....................... ..................................................................... brevicornis BoHEMAN, 1842

– funicular segments globular, segments six and seven slightly transverse; elytra short oval, stout; apex of metatibiae in males almost straight and inconspicuous (6.3–7.2 mm). Gümüshane; Kostandagi Gecidi ...................... ................................................................................ staveni BRAuN, 2000

Figures

Fig. 3. Map providing an overview on all Otiorhynchus (Tecutinus) species described so far: 1) brevicornis BoHEMAN, 1842; 2) caesareus K. DANIEL, 1906; 3) catonii LoNA, 1943; 4) crassicornis GYLLEN- HAL, 1839; 5) cribripennis HocHHuTH, 1851; 6) crinitellus K. DANIEL & Ι. DANIEL, 1902; 7) escherichi REITTER, 1898; 8) fremuthi BRAuN, 1989; 9) gymnopterus K. DANIEL & Ι. DANIEL, 1902; 10) heinzianus BRAuN, 1988; 11) ikisderensis SMREczYÑsKI, 1970; 12) karagolensis SMREczYÑsKI, 1970; 13) kindermanni STIERLIN, 1861; 14) latifrons STIERLIN, 1890; 15) lefkaoriensis GERMANN & CoLoNNELLI, 2015; 16) paracrinitellus BRAuN, 1989; 17) pseudocribripennis BRAuN, 1988; 18) riedeli BRAuN, 1989; 19) salbakosanus LoNA, 1943; 20) soganliensis SMREczYÑsKI, 1977; 21) staveni BRAuN, 2000; 22) tmolosensis LoNA, 1943; 23) torulensis BENEDIKT, 2000; 24) charleshuberi sp. nov. (copyright by Google).