Tomarus villosus ( Burmeister, 1847 )

López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 436-438

Tomarus villosus ( Burmeister, 1847)

( Figs. 35–38 )

Podalgus villosus Burmeister, 1847: 120

Oryctes nitidicollis Solier, 1851: 79

Ligyrus patagonus Steinheil, 1872: 560 Diagnosis. Tomarus villosus can be recognized by the antennal club longer than antennomeres 2–7; clypeus narrowed towards apex (base 2.5 times wider than apex); frontoclypeal tubercles cariniform; pronotal punctation minute and sparse; pronotum without fovea; pronotal tubercle small; males with the inner claw of the protarsi simple;

protibia tridentate, without additional basal denticle; apex of metatibia with 18–25 spinules; parameres with 2 acute ventral teeth, apices straight ( Figs. 36–37 ).

Locality records ( Fig. 38 ). 2 males and 1 female from MEKRB. Arequipa (2): Socabaya, Lara. Lima (1): La Molina .

Temporal distribution. February (2), June (1).

Distribution. Peru, Chile, and Argentina (Endr̂di 1985; Neita-Moreno & Ratcliffe 2017).

Natural history. Gutiérrez (1945) observed the mating behavior of some specimens of T. villosus in Chile, where the males flew searching for females and the copulation occurred below soil, in the same hole from where the females emerged.

Figures

FIGURES 35–38. Tomarus villosus: 35, habitus; 36, parameres in frontal view; 37, parameres in lateral view; 38, locality records in Peru. Habitus photograph by Yony Callohauri.