Exocelina okbapensis hajeki Shaverdo & Balke

Shaverdo, Helena, Sumoked, Bob & Balke, Michael, 2017, Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 715, pp. 17-37 : 23-26

1a. Exocelina okbapensis hajeki Shaverdo & Balke ssp. n. Figs 3, 6

Exocelina undescribed sp. MB0066: Toussaint et al. 2014: Supplementary figs 1-4, tab. 2.

Type locality.

Papua: Jayawijaya Regency, Wamena, 04°03.6'S; 139°01.9'E, 2050 m a.s.l.

Type material.

Holotype: male "IN, PA: Jayawijaya Regen., Baliem vall., 10km NE Wamena, forest above 'Baliem vall. Resort’, 2050 m, 2-3.II.2015, 04°03.6'S, 139°01.9'E; J. Hájek & J. Šumpich leg" (NMPC). Paratypes: 2 males and 7 females with the same label as the holotype, one female with an additional label "M. Balke 7372" (MZB, NHMW, NMPC, ZSM). 6 males, 11 females "INDONESIA, Papua: Jayawijaya Distr., Ballem valley, 10km NE of Wamena, forest above "Baliem valley resort", 04°03.6'S, 139°01.9'E, 2050 m; 2-3.ii.2015 J. Hájek & J. Šumpich leg." (NHMW, NMPC, ZSM). 1 male "66 M. Balke" [green], "measured J. Parkin 43", "Indonesia: Irian Jaya, N Wamena, Cerny, M. Balke: MB 66" (ZSM).

Diagnosis.

Beetle medium-sized; dark brown to piceous, with reddish brown pronotal sides; submatt; pronotum with distinct lateral bead; male antennomeres 2-6 slightly, but evidently enlarged, female antennomeres 2-6 stout; male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, without concavity, with anterior row of 26 and posterior row of eight relatively short setae; median lobe curved, with apex curved downwards and slightly rounded in lateral view. The subspecies differs from the nominative subspecies in the modified antennae, in the shape and setation of the paramere, and in the more striated abdominal ventrite 6; some beetles are also somehow slightly more matt due to more strongly impressed dorsal microreticulation.

Description.

Size and shape: Beetle medium-sized (TL-H 3.75-4.5 mm, TL 4.1-4.9 mm, MW 2.0-2.3 mm), with oblong-oval habitus, broadest at elytral middle (Fig. 3). Coloration: As in nominative subspecies.

Surface sculpture: As in nominative subspecies apart from pronotum and elytra with evident, rather strongly impressed microreticulation, dorsal surface submatt.

Structures: As in nominative subspecies.

Male: Antennomeres 2-6 slightly, but evidently enlarged, antennomere 2 with slightly extended external upper angle (Fig. 3). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 26 and posterior row of eight relatively short setae (Fig. 6D). Median lobe curved, with apex curved downwards and slightly rounded in lateral view and evenly tapering to the slightly truncate apex in ventral view. Paramere with distinct notch on dorsal side and small, evidently separated subdistal part with a tuft of very dense, strong setae; proximal setae inconspicuous (Fig. 6 A–C). Abdominal ventrite 6 with numerous (16-22) long lateral striae on each side.

Holotype: TL-H 4.0 mm, TL 4.4 mm, MW 2.05 mm.

Female: Antennomeres 2-6 stout, in some specimens only slightly more slender than in males, antennomere 2 with slightly extended external upper angle; pro- and mesotarsi not modified and abdominal ventrite 6 without striae.

Variability.

In some males, antennomeres 2-6 more strongly enlarged, in some others less strongly enlarged, similar to those of females.

Distribution.

Papua: Jayawijaya Regency. The subspecies is known only from the type locality (Fig. 12).

Habitat.

Near Wamena, the species was collected from wet ground with weak water flow and forest puddles, which turn to a small creek during rain (Figs 13-14).

Etymology.

The subspecies is named after our friend and colleague Jirí Hájek who collected almost all the specimens. The name is a noun in the genitive case.