Macroderes greeni Kirby, 1818

Abdalla, Ishtiag H., Deschodt, Christian M., Scholtz, Clarke H. & Sole, Catherine L., 2018, An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa, Zootaxa 4504 (1), pp. 41-75 : 56-58

Macroderes greeni Kirby, 1818

( Figs. 10 , 44 , 58 , 91 )

Onthophagus greeni Kirby, 1818: 397 . Macroderes greeni: Westwood 1847: 228 ; Preudhomme de Borre 1880: 9; Péringuey 1901: 298; Janssens 1939: 27; Ferreira 1969: 320; Frolov & Scholtz 2005: 388.

Type locality. Types with no locality data.

Type material examined. None. Type series is deposited in BMNH ( Frolov & Scholtz 2005).

Additional material examined. 1 ♂ (TMSA): “S.Afr / SW Cape Prov / Abrahamskraal farm / 33°.14' S– 18°.09' E // 25.8.1983 / E–Y: 1976 / groundtraps / 70 days / Endrody, Penrith leg // groundtraps with faeces bait // Macroderes greeni (Kirby) / det. A. Frolov / 2003”; 3 ♀♀ (TMSA): “S.Afr / S.W Cape / Struisbaai / 34°.46' S– 20°.03' E // 28. 8. 1983 // E–Y: 1987 / groundtrap / 60 days / Endrody, Penrith leg // groundtraps with faeces bait // Macroderes greeni (Kirby) / det. A. Frolov / 2003”; 1 ♂ and 7 ♀♀ (SANC): “LANGEBAAN / C P / 12 Km / SE / Farm Geelbek / A.L.V. Davis / Macroderes sp / det. CSIRO. DBRU // Macroderes greeni (Kirby) / Frolov / det / 2003 but with different dates of collecting: 1♀: 6.VII.1979; 1 ♂ and 2 ♀♀: 13.VIII.1979; 1 ♀: 23.V.1979; 1 ♀: 30.V.1979. 1 ♀: 1–29. X. 79; 1 ♀: 26. VI. 1979; 2 ♂♂ and 4 ♀♀ (SANC): “S.Afr / SW / CP / Modderrivier farm / nr Atlantis / 33° 28' S 18 20' E / 29°. VI.1987 / ALV. Davis // ex cattle dung baited / pitfall in Strandveld // Macroderes greeni (Kirby) / Frolov / det / 2003”; 1 ♀ (SAMC): “ Macroderes greeni // Macroderes greeni (Kirby) / det. Frolov / 2003”; 1 ♂ (SAMC): “Saldanha Bay // greeni Kirby // Macroderes greeni Kirby / det. Frolov / 2003 // SAM—A 043223”; 2 ♂♂ (SAMC): “Mamre-Mamlesburg Rd / Sep. 1938 // Macroderes greeni Kirby / det. Frolov / 2003 // SAM-Col-A02321”.

Size range. Males length: 9.6–11.6 mm, width: 6.2–8.0 mm; females length: 10.3–11.4 mm, width: 6.3–7.8 mm.

Differential diagnosis. This species shares the elytral punctuation shape of M. arrowi , M. foveatus and M. cornutus ( Fig. 91 ), however, it can differentiated by the large and rounded punctures of the pronotum ( Figs. 44 , 58 ) and from the latter two species by lacking the triangular concavity in the base of the pronotum which characterise the two species.

Habitat and distribution. This species occupies a small range which extends along the south-western coast of South Africa from Cape Agulhas in the southeast and up to Saldanha Bay in the northwest ( Fig. 2). The area is situated within the Southwest Fynbos Bioregion which is considered as the core of Fynbos Biome ( Mucina & Rutherford 2006). The area is mostly sandstone typified by winter rains and low to very low summer rains.

Figures

FIGURES 7–10. Habitus of Macroderes species. 7, M. fornicatus Sharp, 1880; 8, M. foveatus Frolov & Scholtz, 2005; 9, M. bias (Olivier, 1789); 10, M. greeni (Kirby, 1818).

FIGURES 40–45. Pronotum of Macroderes species. 40, M. foveatus Frolov & Scholtz, 2005; 41, M. cornutus Frolov & Scholtz, 2005; 42, M. namakwanus Frolov & Scholtz, 2005; 43, M. politulus Preudhomme de Borre, 1880; 44, M. greeni (Kirby, 1818); 45, M. endroedyi Frolov & Scholtz, 2005.

FIGURES 58–63. Punctuation of the base of pronotum of Macroderes species. 58, M. greeni (Kirby, 1818); 59, M. arrowi Janssens, 1939; 60, M. bias (Olivier, 1789); 61, M. namakwanus Frolov & Scholtz, 2005; 62, M. endroedyi Frolov & Scholtz, 2005; 63, M. politulus Preudhomme de Borre, 1880.

FIGURES 88–93. Pronotum punctures shape (88, 89) and sculpture of second elytra intervals (90–93) of Macroderes species. 88, M. soleiana Abdalla & Deschodt, new species; 89, M. tourtosus Abdalla and Scholtz, new species; 90, M. minutus Frolov and Scholtz, 2005; 91, M. greeni (Kirby, 1818); 92, M. politulus Preudhomme de Borre, 1880; 93, M. gifboomi Abdalla & Scholtz, new species.