Raveniola shangrila, Zonstein, Sergei & Marusik, Yuri M., 2012

Zonstein, Sergei & Marusik, Yuri M., 2012, A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan, ZooKeys 211, pp. 71-99 : 77-79

Raveniola shangrila sp. n. Figs 1119273341

Types.

Holotype ♂ - CHINA: Yunnan Province, Sueshan Mt. Ridge, Shika Mts. 10-15 km W Zhongdian (approximately 27°48'N, 99°35'E), 3800-4300 m, 25.05- 6.06.2005, coll. I. Shokhin & S. Murzin (IZAS). Paratypes. 5♂ with the same collecting data are shared between MNHG, MNHN, NHML, TAU and ZMMU.

Etymology.

The specific epithet is given in honour of the mythical Tibetan land Shangri-La attributed to the highland region located in the far eastern part of Tibet (Xizang) and north-western part of Yunnan, i.e., including the type locality of this species.

Diagnosis.

The full reduction of PMS allows to place this species together with Raveniola guangxi ; Raveniola shangrila can be distinguished from the latter species by shape of the embolus and larger number of maxillary cuspules - 15-20 vs. 3-4 (cf. Figs 15, 30, 37 and 19, 33, 41, respectively). In general, specimens of Raveniola shangrila sp. n. appear poorly distinguishable from those of Raveniola songi sp. n., but certain distinctions in the configuration of male bulb and metatarsus I are evident (cf. Figs 15, 16, 23 and 24).

Description.

Male (holotype). Body length 16.10. Colour in alcohol: carapace (with lighter spotted pattern), legs I partially, legs II–IV mostly middle foxy brown; sternum, labium and maxillae lighter coloured; chelicerae, all femora dorsally, tibiae and metatarsi I dark reddish brown; eye tubercle blackish brown; abdomen uniformly light brownish grey; genital area, booklungs and spinnerets pale yellowish grey.

Carapace (Fig. 11) 5.91 long, 5.45 wide; covered with semi-adpressed dark hairs. Eye diameters (AME, ALE, PLE, PME): 0.20(0.25), 0.31, 0.20, 0.20. Interdistances: AME–AME 0.15(0.11), ALE–AME 0.12(0.10), ALE–PLE 0.09, PLE–PME 0.03, PME–PME 0.57. Cheliceral furrow with 9 promarginal teeth and 4-5 mesobasal denticles. Labium (Fig. 19) 0.60 long, 1.03 wide. Maxillae with 17-19 small cuspules in wide triangle area. Sternum 2.85 long, 2.75 wide. Palp: 9.19 (3.56, 1.78, 2.77, -, 1.08). Leg I: 18.61 (5.02, 2.85, 4.09, 4.27, 2.38). Leg II: 16.11 (4.65, 2.56, 3.25, 3.46, 2.19). Leg III: 13.48 (3.71, 1.97, 2.47, 3.16, 2.17). Leg IV: 17.57 (4.72, 2.40, 3.81, 4.23, 2.41). Leg I: tibia incrassate, metatarsus curved retroventrally (Fig. 27).

Spination. All femora with a few stiff bristles (undeveloped spines) located medially and distally; patella IV and tarsi I–IV aspinose. Palp: femur pd1; patella p1; tibia d1 –1– 1, p1 –1– 1, r0 –1– 1, v1 –2– 0; cymbium d4. Leg I: femur pd1 –0– 1; patella p1; tibia p1 –1– 1, v2 –2–M–M; metatarsus rv1. Leg II: femur pd1 –0– 1, rv0 –0– 1; tibia p1 –1– 1, v2 –2– 2; metatarsus p0 –1– 1; v2 –2– 3. Leg III: femur pd1 –0– 1, rd1 –0– 1; patella p1-1, r1-1; tibia d1-0, p1-1, r0-1, v2 –2– 2; metatarsus d0 –1– 1, p1 –1– 1, r0 –1– 1, v2 –2– 2(3). Leg IV: femur pd0 –0– 1, rd0 –0– 1; tibia p0(1) –1– 1, r0 –0(1)– 1, v2 –2– 2; metatarsus p0 –1– 1, r0 –1– 1, v2 –2– 2.

Scopula: long, distal 1/2 on metatarsus I and II, entire on tarsi I and II, mixed with setae on tarsi III and IV. Paired claws with 6-8 teeth on promargin and retromargin. Trichobothria: 2 rows of 7-9 per row on tibiae, 12-15 on metatarsi, 10-12 on tarsi, 7 on cymbium.

Cymbium with few rather short spines (Fig. 33). Bulb without ridges; embolus long and twisted (Fig. 41).

Spinnerets. PMS: absent. PLS: maximum diameter 0.55; length of basal, medial and apical segments 0.84, 0.54, 0.75; total length 2.13; apical segment digi tiform.

Female unknown.

Variability.

Carapace length in males varies from 5.03 to 5.95 (n=5).

Distribution.

CHINA: Yunnan Province (Fig. 1).