Anchista brunnea (Wiedemann, 1823)

Shi, Hongliang, Zhou, Hongzhang & Liang, Hongbin, 2013, Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera, ZooKeys 284, pp. 1-129 : 22-24

Anchista brunnea (Wiedemann, 1823) Habitus: Figs 4 37, 38 male genitalia: Figs 70 97

Anchista brunnea Wiedemann 1823: 59 (original: Lebia ; type locality: Bengalia; lectotype deposited in ZMUC); Andrewes 1921: 172 ( Anchista ); Andrewes 1930d: 22 ( Anchista , catalogue); Csiki 1932: 1456 ( Anchista , catalogue).

Anchista modesta Nietner 1856: 523 (type locality: Ceylon; syntype deposited in MZPW); Nietner 1857: 375 ( Anchista , redundant publication); Chaudoir 1877: 239 ( Anchista ); Andrewes 1924a: 140 ( Anchista , synonymized with brunnea Wiedemann); Andrewes 1927b: 103 ( Anchista ). [Synonym]

Anchista picea Chaudoir 1877: 238 (type locality: Deccan; lectotype deposited in MNHN); Andrewes 1921: 172 ( Anchista , synonymized with brunnea Wiedemann). [Synonym]

Type examined.

Lectotype of Lebia brunnea Wiedemann, designated herein (ZMUC): female, body length = 7.4 mm, pin mounted, "Mus. / Westerm."; “Type” [red label]; " Bengal. / Mai 1809. / D: brunneus. /Lebia brun / nea Wied. "; "LECTOTYPE ♀/ Lebia brunnea / Wiedemann, 1823 / Des. SHI H. L. 2011" [red label][Fig. 37]. Lectotype of Anchista picea Chaudoir, designated herein (MNHN), male, body length = 7.2 mm, pin mounted, "Ex Musaeo, Chaudoir" [red letter]; “TYPE” [red label]; "MUSEUM PARIS / 1952 / COLL. R. OBERTHÜR”; " picea Chaudoir, Deccan, Stevens " [large box label but pinned under specimen]; "LECTOTYPE ♂/ Anchista picea / Chaudoir, 1877 / Des. SHI H. L. 2011" [red label][Fig. 38]. Paralectotype of Anchista picea Chaudoir (MNHN): 4 males, "Ex Musaeo, Chaudoir" [red letter]; “PARATYPE” [red label]; "MUSEUM PARIS / 1952 / COLL. R. OBERTHÜR”; "PARALECTOTYPE ♂/ Anchista picea / Chaudoir, 1877 / Des. SHI H. L. 2011" [red label] [Figs 4, 70, 97].

Notes on types.

Lebia brunnea Wiedemann: In the original literature, Wiedemann didn’t mention how many specimens belonged to the type series. We borrowed one female ( Fig. 37 ) from ZMUC with a red label “Type” and Wiedemann’s hand-written label. We herein designate this specimen as lectotype for taxonomic purpose of fixing the name to unique name-bearing type.

Anchista picea Chaudoir: Five specimens were mentioned in the original description. In the collection of MNHN, these specimens used to be in Chaudoir’s box, but were moved to Lebiini sorted boxes by J. Mateu. Obviously, Mateu selected the first one, and put Chaudoir’s box label and a red “type” label under it. We herein designate this specimen ( Fig. 38 ) with such labels as lectotype for this species for taxonomic purpose of fixing the name to unique name-bearing type.

Non-type material examined (Total 7 specimens from India). 1 specimen (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, XI-XII 1896". 1 specimen (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, VIII-IX 1896". 2 specimens (NHML), "India Nevinson Coll. 1918-14.". 1 specimen (NHML), "Fyzabad, Unt. Prov., India. R. W. G. Hingston."; "Anchista brunnea Wied (= picea Chaud.), H. E. Andrewes det.". 1 specimen (NHML), "Nagpur., C. P. India, 1000ft."; "Anchista brunnea Wied, = picea Chaud., Compared with 2 types H. E. A.". 1 female (NMPC), "Dabhalwala, Dehra Dun, India".

Diagnosis.

Head and pronotum glabrous; pronotum widest at middle, lateral margins slightly angulate in middle; elytra uniform brownish; 3rd interval of elytra with two pores, 5th interval with one pore near base.

Based on the glabrous surface and number of setigerous pores on 3rd and 5th intervals, this species can be readily distinguished from all the other species of Anchista . Other species with the same pore number all belong to different genera ( Paraphaea , Metallanchista gen. n.). They are different in elytral pattern and pronotum shape and have other generic differences.

Description.

Body length 7.2-7.8 mm; head and pronotum reddish brown; antennae and mouthparts yellowish brown or reddish brown, 1st antennomere and apices of terminal maxillary and labial palpomeres slightly paler; pronotum with lateral explanate area yellowish brown; elytra dark brown, without distinct pattern, but usually with basal two-thirds of disc slightly paler; ventral side same color as dorsal side; femora yellow, tarsi and tibiae yellowish brown. Head glabrous, without punc tures or microsculpture. Pronotum glabrous,cordiform, widest slightly before middle; ratio PW/PL 1.38 to 1.41; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles widened and rounded, not projected, with a few short setae; lateral margins slightly angulate in middle, strongly sinuate before hind angles; lateral explanate areas wide and even, with a few coarse punctures; hind angles sharp, rectangular or slightly acute, with a few very short accessory setae; basal foveae moderately deep, with a few punctures; median line distinct, strongly depressed and punctate anteriorly and subbasally. Elytra with striae shallow, with fine punctures; intervals hardly convex, without accessory setae, sparsely and finely punctate; microsculpture distinct, isodiametric; 3rd interval with two setigerous pores, basal one placed at basal one-third approximately, adjacent to 3rd interval, apical one placed at apical one-eighth approximately, adjacent to 2nd stria; 5th interval with one setigerous pore near base, adjacent to 5th stria; basal margination nearly complete. Males with adhesive hairs well developed (two whole rows) on 1st to 3rd mesotarsomeres. Male genitalia with median lobe of aedeagus straight and flat, slightly expanded to apex; left-lateral margin curved in middle; apical orifice opened dorsally, basal margin of apical orifice setose; apical lamella placed on right side, flat, short and rounded in dorsal view, slightly curved upward dorsally in lateral view; internal sac with two elongate and sinuate weakly sclerotized sclerites near apical orifice, basal part of internal sac finely scaled ( Fig. 70 ). Female genitalia not studied.

Distribution

( Map 3 ). India, Bengal, Sri Lanka.

Notes on synonym.

Andrewes (1924a, 1927b) indicated that Anchista modesta Nietner is a synonym of Lebia brunnea Wiedemann. We didn’t examine the type of Anchista modesta , which should be in the Warsaw Museum, nor any material from Sri Lanka. We follow Andrewes’ treatment in the present paper.

Figures

Map 3. Known distribution of genus Anchista: ◆ Anchista brunnea (Wiedemann) ★ Anchista nubila Andrewes ● Anchista pilosa sp. n. △ Anchista fenestrata fenestrata (Schmidt-Goebel) ▼ Anchista fenestrata subpubescens Chaudoir.

Figure 70. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir.

Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal.

Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes.

Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian.