Pupina billeti Fischer, 1898

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina billeti Fischer, 1898

Figs 21S , 26C

Pupina billeti Fischer, 1898: 333, 334, pl. 18, figs 38-41. Type locality: Rochers calcaires Déo-Ma-Phuc [limestone areas around Ma Phuc Pass, Tra Linh District, Cao Bang Province, Vietnam]. Fischer and Dautzenberg 1904: 431, Bac-Kan, Tonkin [Bac Kan Province, Vietnam].

Pupina (Tylotoechus) billeti - Kobelt 1902: 309.

Type material examined.

Holotype MNHN-IM-2000-35841 (Figs 21S , 26C ) from Deo-Ma-Phuc.

Diagnosis.

Shell ovate; last whorl ca. 70% of shell height. Apertural lip extremely thickened; with a furrow between inner and outer peristomes; inner peristome thickened and cord-like; parietal callus distinct. Both parietal and columellar teeth extremely thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.

Differential diagnosis.

Pupina billeti can be distinguished from all other species in the P. artata species group from mainland Southeast Asia by having the thickest parietal and columellar teeth and apertural lip, and a distinct parietal callus.

Distribution.

Northern Vietnam ( Fischer and Dautzenberg 1904).

Remarks.

As P. billeti was described based on a single specimen as explicitly stated in the original description, that specimen is the holotype fixed by monotypy ( ICZN 1999: Art. 73.1.2).

Figures

Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T).

Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D).