Drymeia ponti, Savage & Sorokina, 2021

Savage, Jade & Sorokina, Vera S., 2021, Review of the North American fauna of Drymeia Meigen (Diptera, Muscidae) and evaluation of DNA barcodes for species-level identification in the genus, ZooKeys 1024, pp. 31-89 : 31

Drymeia ponti sp. nov. Figs 1A , 2A , 10A , 11D , 13 , 14 , 15

Type material.

Holotype male labelled "W. Side Cortes Pass/ 1100' Mexico, Mexico / 13-VIII-1954 / J.G. Chillcott"; " HOLOTYPE / Drymeia ponti ♂ / Savage & Sorokina [red]" ( CNC). Paratypes : all with " PARATYPE / Drymeia ponti / Savage & Sorokina [yellow]" : 2 males and 15 females, same as holotype. 1 male and 2 females, same as holotype except ( BUIC) . 1 male and 5 females labelled " Rio Frio / Fed. Dist. Of Mex. / I.IX.69/ D. Kritsch " ( CNC) . 1 male, same as previous except ( BUIC) . 1 male labelled " Rio Frio , MEX./ MWX.,27-VIII-69/ D. Kritsch BL 10,000'" ( CNC) .

Etymology.

The species name is a patronym in honour of Adrian C. Pont ( UK), an exceptional dipterist and mentor to both co-authors.

Diagnosis.

Small dark species with strong prealar, 2+3 dc and strong costal spine. This species is similar to Drymeia aterrima (Wulp, 1896), especially in the dark male calypter, but can be distinguished from it in the male by a broad frontal vitta (Fig. 13B ) and a distinct ventral apical process on T 3 (Fig. 13D ), and in the female by the presence of a flattened fore tarsomere 5 (Fig. 15C ).

Description.

Male. Body length: 4.1-5.7 mm; wing length: 3.8-5.2 mm.

Head: Ground colour black; eye bare; fronto-orbital plate and parafacial dark brown pruinose; face black, gena and lower occiput dark brown pruinose; frons at narrowest point approximately 1.5 × width of ocellar triangle with black frontal vitta exposed (Fig. 13B ); parafacial in lateral view equal to or slightly wider than width of first flagellomere along most of its length (Fig. 13A ); lower margin of the face projecting slightly beyond lower level of frons in lateral view; gena at narrowest point as high as length of first flagellomere, densely setose and without a group of upcurved setae on anterior part of genal dilation; 9-12 frontal setae (including interstitials) reaching to anterior ocellus; antenna black; first flagellomere 1.2 × as long as wide; arista swollen near base and pubescent, with longest hair as long as basal diameter of arista; palpus black; proboscis long and narrow with prementum approximately 2 × as long as palpus, undusted and glossy; labella small.

Thorax: Ground colour black; scutum, postpronotum, notopleuron, postalar callus and scutellum with dense brown dusting; pleuron dark brown, slightly shiny with brown dusting; katepisternum with small anterior undusted glossy patch, meron entirely dusted; anepimeron and katepimeron bare; notopleuron setulose; acr 0+1; 2+3 dc; prealar long and strong, as long as second notopleural.

Legs: Black; T 1 with 2 pv on apical 1/2; F2 straight, with matching rows of long strong av and pv on apical 2/3, these much longer than width of femur (Fig. 13A, C ), a complete row of short ad, and 2 or 3 preapical pd-p; T 2 with 1 long and 2 shorter ad on apical 1/2, 4 pd and 1 pv; F3 with a complete row of ad, a row of av, longer on apical 1/2, a row of p on basal 2/3 and a row of long strong pv on apical 1/3; T 3 with 2 av, 4 ad, 3 pd and with 4 short delicate pv on apical 1/2, ventral apical process short but distinct (Fig. 13D ), apical pv absent or reduced, no longer than 1/2 the length of apical av when visible.

Wing: Brown, darker at base with dark brown veins; basicosta and tegula black; costal spinules strong, with costal spine 2 × as long as costal spinules; calypters with membrane and edges dark brown.

Abdomen: Conical; ground colour black; brown dusted, subshiny with no median vittae; sternite I bare; sternite V as in Fig. 14C .

Terminalia: Fig. 14A, B .

Female. Body length: 4.5-6.5 mm; wing length: 4.0- 5.5 mm. Differs from the male as follows:

Head: Frontal triangle undefined; frontal vitta black; parafacial mostly dusted, with a small narrow shiny patch near base of antenna (Fig. 15A, B ); frons at midpoint approximately 0.4 × as wide as head and approximately 0.9 × as long as wide; fronto-orbital plate narrow, approximately as wide as distance between inner margins of posterior ocelli; three or four medioclinate frontal setae and several weaker interstitials, three orbital setae, the upper two reclinate and lateroclinate, the lower one proclinate.

Thorax: As in male.

Legs (chaetotaxy described in full): T 1 with 1 or 2 pv; fore tarsomere 5 distinctively flattened (Fig. 10A , 15C ); F2 with 1 or 2 prebasal and 2 or 3 preapical av without pv; T 2 with 0 or 1 av (most without), 2 or 3 ad, 4 or 5 pd and 2 pv; F3 with av row complete, without pv; T 3 with 2 or 3 av, 3 or 4 ad and 3 pd, apical pv usually absent but if present, then no longer than 1/2 length of apical av.

Wing: Light brown, darker near base; with membrane and edges dark yellow.

Abdomen: as in male.

Distribution.

Neotropical: Mexico ( Mexico).

DNA Barcode.

None available.

Figures

Figure 1. Male frons, frontal view A D. ponti B D. aterrima C D. neoborealis D D. vockerothi. Scale bars: 0.5 mm.

Figure 2. Male hind tibia, dorsal view A D. ponti B D. aterrima C D. groenlandica D D. setibasis E D. similis F D. vockerothi G D. quadrisetosa H D. glacialis I D. hucketti J D. woodorum. Scale bars: 0.5 mm.

Figure 10. Female fore tarsomeres 3 - 5 A D. ponti B D. aldrichi C D. similis D D. profrontalis E D. minor. Scale bars: 0.25 mm.

Figure 11. Female frons A D. similis B D. latifrons C D. profrontalis D D. ponti E D. hucketti. Scale bars: 0.5 mm.

Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D).

Figure 14. Drymeia ponti sp. nov., male terminalia A lateral view B posterior view C sternite V.

Figure 15. Drymeia ponti sp. nov. A female habitus, paratype B female head, frontal view C female tarsomeres 3 - 5. Scale bars: 1 mm (A, B); 0.5 mm (C).