Ummidia asperula (Simon, 1889)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia asperula (Simon, 1889) Figs 68 , 69 , Map 6

Pachylomerus asperulus Simon, 1889: 179; HOLOTYPE 1 ♀ (AR4147) from Venezuela (label); from Catia La Mar near Caracas, north slopes of Silla near Maiquetía, Venezuela, 10.5778 -66.9973 5, 261 m a.s.l. (Simon, 1889), deposited in MNHN, examined.

Diagnosis.

Ummidia asperula is a relatively small species which can be differentiated from all geographically proximate species by the lack of a distinct tarsal comb on the retrolateral face of tarsus IV in both males and females and from U. erema , U. tibacuy , and U. neblina by tarsi not being light in color. Males can be further distinguished from U. erema by the embolus being of moderate length with a single even curve, by having few prolateral spines on tibia I (3 vs 15). Females can be differentiated from all geographically proximate species except U. salebrosa by having narrow stalked spermathecae which curve slightly medially and then coil with bulbs facing dorsally. Females can be distinguished from U. tunapuna by having all eyes on a high, defined tubercle and by lacking a setal fringe on the lateral and posterior edges of the carapace.

Description of male exemplar.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp and leg I, right leg I removed, in vial with specimen. General coloration. Carapace and chelicerae reddish black 2.5YR 2.5/1, legs very dusky red 2.5YR 2.5/2, tarsi strong brown 7.5YR 5/8. Abdomen dark reddish gray 2.5YR 3/1. Cephalothorax. Carapace 3.69 long, 3.66 wide. Pars cephalica 2.62 long. Foveal groove procurved, 0.32 long, 0.69 wide. Tubercle high. AER procurved. PER slightly procurved. Eye group 0.6 long, 1.07 wide, AME 0.34, PME 0.18, ALE 0.37, PLE 0.21. Sternum sparsely setose around outer 1/3, STRl 2.05, STRw 1.96. Chelicerae with anterior tooth row comprising five teeth, posterior margin with six teeth. Palpal endites with 16 small subhastate cuspules spread across proximal half of endite face, lacking distal endite cuspules, ENDw 0.73, ENDl 1.41. Labium with nine cuspules, LBw 0.77, LBl 0.58. Rastellum with eight spines along distal cheliceral margin, three on process. Abdomen setose. Legs. F1 3.71; F1w 0.78; P1 1.61; Ti1 2.56; Mt1 1.65; Tr1 0.93; F3 2.64; F3w 0.91; P3 1.28; Ti3 1.56; Sd3 1.1; Mt3 1.33; Tr3 1.12; F4 3.37; F4w 0.88; P4 1.44; Ti4 1.97; Mt4 1.91; Tr4 1.05. Retrolateral face of tarsus IV without brush or comb. Leg I spination pattern: TSp 1, TSpv 2, TSrd 0, TSr 3, TSrv 23, MtSp 0, MtSr 8, TrSp 0, TrSr 1. Pedipalps. PTl 1.86, PTw 0.71, Bl 1.39. Embolus evenly curved.

Variation, males.

Known only from male exemplar.

Description of female holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. Left leg III removed, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4. Abdomen very dark brown 7.5YR 2.5/2, spinnerets yellowish brown 10YR 5/6. Cephalothorax. Carapace 7.61 long, 7.61 wide. Pars cephalica 5.75 long. Foveal groove procurved, 0.86 long, 1.7 wide. Eye tubercle high, defined, and with all eyes on and close set. AER procurved. PER procurved. Eye group 1.01 long, 1.39 wide, AME 0.34, PME 0.32, ALE 0.55, PLE 0.39. Sternum moderately setose, STRl 4.8, STRw 4.48. Chelicerae with anterior row comprising six teeth, posterior margin with seven teeth. Palpal endites with 21 cuspules spread across proximal half of endite and 19 cuspules distally, ENDw 1.86, ENDl 3.27. Labium with nine cuspules, LBw 1.75, LBl 1.24. Rastellum with many strong sines on process with heavy hairs on cheliceral face above process. Abdomen. Evenly setose with pale speckles concentrated at apodemes and slight lightening along the dorsal midline. Legs. F1 5.05; F1w 2; P1 3.3; Ti1 3.06; Mt1 2.01; Tr1 1.04; F3 4.48; F3w 2.35; P3 2.91; Ti3 2.75, Sd3 1.71; Mt3 1.82; Tr3 1.46; F4 5.44; F4w 2.24; P4 3.39; Ti4 3.2; Mt4 3.09; Tr4 1.64. Retrolateral face tarsus IV with loose comb of spinules surrounded in hairs. Pedipalps. PF 4.71, PP 2.9, PTi 2.85, PTr 2.52. Spermathecae with slight medial bend, small bulbs facing dorsally.

Variation, females.

Known only from female holotype.

Material examined.

Venezuela: Catia La Mar near Caracas, north slopes of Silla near Maiquetía, 10.5778 -66.9973 5, 261 m a.s.l. (AR4147, 1889, 1♀, Simon , MNHN) Bolivar: Rancho Grande , 6.4142 -66.5169 9, 188 m a.s.l. (UMM577, 13.v.1954, 1♂, W Beebe et. al., AMNH) .

Figures

Map 6. Locality records for South American and Caribbean species.

Figure 68. Ummidia asperula (Simon 1889) from Venezuela A male habitus illustration UMM 577 B female habitus illustration AR 4147. Scale bars: 4.0 mm.

Figure 69. Ummidia asperula (Simon 1889) from Venezuela A-D male exemplar (UMM 577) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae AR 4147. Scale bars: 1.0 mm.