Ceratozamia kuesteriana Regel, Bull. Soc. Imp. Naturalistes Moscou 30: 187. t. 3. 1857
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https://dx.doi.org/10.3897/phytokeys.208.80382 |
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https://treatment.plazi.org/id/AD3592D8-91D2-5D3C-B414-A71FEFB90639 |
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scientific name |
Ceratozamia kuesteriana Regel, Bull. Soc. Imp. Naturalistes Moscou 30: 187. t. 3. 1857 |
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14. Ceratozamia kuesteriana Regel, Bull. Soc. Imp. Naturalistes Moscou 30: 187. t. 3. 1857
Figs 3D View Figure 3 , 7A View Figure 7 , 9B View Figure 9
Type.
Cultivated in St. Petersburg, Russia "Ex Horto Petropolitano", 1856, E. Regel s.n. (holotype: LE! [LE00009046]; isotype: U! [U0007272]) .
Description.
Stem 10-30 cm long, 10-25 cm in diameter, semi-hypogeous, erect. Cataphylls 1.5-4 × 2-3.5 cm wide at the base, persistent, triangular, reddish brown, densely brown tomentose at emergence, partially tomentose at maturity, apex acuminate. Leaves 1-11, 80-133 cm long, ascending, reddish brown at emergence, with whitish gray trichomes, glabrous at maturity. Petiole 30-72 cm long, terete, linear, greenish brown in mature leaves; with 1-10 thin prickles, 0.01-0.15 cm long. Rachis 40-72 cm long, terete, linear, greenish brown in mature leaves, with prickles and rarely unarmed. Leaflets 22-50 pairs, opposite to subopposite, insertion in one plane, linear, adaxially curved, basally falcate, papyraceous, caniculate, green, adaxial and abaxial sides glabrous, distal end with entire margins, acuminate and symmetrical at the apex, attenuate at base, with conspicuous and light veins; median leaflets 17-32 × 0.6-1 cm, 0.3-2.5 cm between leaflets; articulations 0.2-0.8 cm wide, brown. Pollen strobili 11-15 cm long, 1.6-2.5 cm in diameter, solitary, cylindrical, erect, greenish yellow with brown trichomes at emergence, yellowish brown with reddish brown trichomes at maturity; peduncle 8-14 cm long, 0.7-1 cm in diameter, reddish brown to brown pubescent; microsporophylls 0.6-1.1 × 0.5-1 cm, discoid with a non-recurved distal face and a lobate fertile portion, infertile portion 0.28-0.32 cm long and linear with straight horns 0.05-0.25 cm long, 0.45-0.53 cm and an acute angle between the horns. Ovulate strobili 13-21 cm long, 7-9 cm in diameter, solitary, cylindrical, erect, green at emergence with brown trichomes, grayish light green with black trichomes at maturity, acuminate apex; peduncle 9-16 cm long, 1-2 cm in diameter, erect, tomentose, brown; megasporophylls 30-72, 6-8 orthostichies with 5-9 sporophylls per orthostichy, 2.5-4.0 × 2.3-3.5 cm, with a truncate distal face, horns straight and 0.30-0.52 cm long, 0.60-0.94 cm between horns with an obtuse angle between the horns. Seeds 1.5-2.3 cm long, 1.2-1.8 cm in diameter, ovate, sarcotesta whitish red when immature, light brown at maturity.
Distribution and habitat.
Ceratozamia kuesteriana is endemic to Tamaulipas State, Mexico (Fig. 19B View Figure 19 ), where it occurs on karstic rocks in pine-oak and cloud forests at 1,100-1,500 m.
Etymology.
The specific epithet is in honor of Baron K. von Kuester.
Common names.
None recorded.
Uses.
None recorded.
Conservation status.
( IUCN 2021). The status for Ceratozamia kuesteriana "Critically Endangered" under criteria A2cd; B2ab(ii,iii,iv).
Discussion.
Ceratozamia kuesteriana is similar to C. sabatoi , but it is easily distinguished by its canaliculate leaflets and ascending leaves.
Specimens examined:
Mexico. Tamaulipas: Mun. Gómez Farías , 1,260 m, 1 May 1967, A. Gómez-Pompa 2029 (MEXU); 1,150 m, 12 Aug 1983, A.P. Vovides 791, 800, 2056 (XAL); 13 Aug 1983, A.P. Vovides & G. Pattison 771, 772, 801 (XAL); 27 Jun 2017, C.I. Carvajal-Hernández et al. 1355 (CIB); 1,200-1400 m, 20 Nov 1970, F. González-Medrano & E. Martínez 3288 (MEXU); 1,100 m, 21 Jan 1970, F. González-Medrano et al. 3362 (MEXU, MO); 13 Aug 1985, L. Trejo s/n (UAT); 1,050 m, 30 Mar 1961, P.S. Martin & C. Saravia 1170 (ENCB). Mun. Ocampo, 1,255 m, 14 Jan 2001, T.W. Walters TW-201-11-A (XAL), TW-201-11-B (MEXU). Mun. Tula, 1,450 m, 21 Jul 1983, D.W. Stevenson 569H, 569G, 569K (MEXU); 1,296 m, 19 Mar 2016, F. Nicolalde-Morejón et al. 2357 - 2365 (CIB); 1,296 m, 19 Mar 2016, L. Martínez-Domínguez et al. 854 - 867, 869 (CIB), 868 (CIB, MEXU); 1,140 m, 19 Mar 2016, L. Martínez-Domínguez et al. 870 - 881 (CIB); 24 Apr 2001, S. Avendaño 5328 (MEXU).
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