Malcolmia aculeolata (Boiss.) Boiss., Fl.

AL-Shehbaz, Ihsan A. & Barriera, Gabrielle, 2019, Typification of Edmond Boissier’s Cruciferae (Brassicaceae) names enumerated in Flora Orientalis, Boissiera 72, pp. 1-193 : 66

Malcolmia aculeolata (Boiss.) Boiss., Fl. Orient. 1: 226. 1867.

 Sisymbrium aculeolatum Boiss. in Ann. Sci. Nat., Bot. ser. 2, 17: 75. 1842.

Type: “[Aucher-Eloy] Ispahan absque numero”.

Holotypus: IRAN: “ Ispahan ”, s.d., Aucher-Eloy s.n. ( P [ P05384929 ]; iso-: W [ W18890303536 ]) .

Notes. – There is no material in the Geneva herbaria of the species with Aucher-Eloy s.n. It is certain that BOISSIER (1842a) based his description on the unicate P 05384929 that he annotated, and he did not examine the duplicate at W. Unfortunately, BOISSIER (1867a, 1888) never mentioned the type collection of the species. HEDGE (1968: 325) was correct in stating that the type is Aucher-Eloy s.n., though he did not indicate the herbarium housing that collection. However, the listing of the type as Schimper 124 from Sinai by JAFRI (1973) is erroneous, and that collection was not cited in the original publication though listed, among others, in Flora Orientalis.

The initial placement of the species in Sisymbrium by BOISSIER (1842a) is currently accepted both on morphological ( WARWICK & AL-SHEHBAZ, 2003) and molecular grounds ( WARWICK et al., 2002). Although, the species was transferred by him to Malcolmia , its lack of the branched trichomes and decurrent stigmas stands against such assignments. Furthermore, the placement in Torularia by HEDGE (1968) or Neotorularia by Léonard and Hedge (in LEONARD, 1986) is not acceptable because it has simple instead of the branched trichomes characteristic of the last genus.