Heerz macrophthalma Martinez , Zaldivar-Riveron , Ceccarelli & Shaw

Zaldivar-Riveron, Alejandro, Martinez, Juan Jose, Ceccarelli, Fadia Sara & Shaw, Scott R., 2012, Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico, ZooKeys 164, pp. 1-23 : 8-10

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Heerz macrophthalma Martinez , Zaldivar-Riveron , Ceccarelli & Shaw

sp. n.

Heerz macrophthalma Martinez, Zaldivar-Riveron, Ceccarelli & Shaw   ZBK   sp. n. Figs 3 A–D


Heerz macrophthalma   distinguishes from the remaining species of the genus by its considerably larger eyes (Fig. 3B). It can also be distinguished from the Central and South American species, Heerz tooya   and Heerz lukenatcha   , by its entirely hyaline wings (Fig. 3A) [wings partially or totally infuscate in the latter two species (Fig. 2B)] and from Heerz ecmahla   by the striate sculpture on the second metasomal tergite (Fig. 3D) [coriaceous in Heerz ecmahla   (Fig. 2C)].


Female. Colour: Body mostly honey yellow, antennae honey yellow, turning darker apically; legs creamish white, with apex of tarsomeres darker; wings hyaline, veins, pterostigma and tegula light brown. Body length: 4.7 mm. Head: vertex, frons and temple striate, face acinose; eyes large, 1.3 times higher than wide (lateral view); malar space about 0.2 times eye height; ocello-ocular distance less than 0.8 times diameter of lateral ocellus and as long as posterior ocellar line; antennae with 31 flagellomeres. Mesosoma: length of mesosoma 1.8 times its maximum height; pronotum smooth, pronotal groove scrobiculate; mesoscutal lobes coriaceous; notauli deep and scrobiculate, obscured in a posterior striate-rugose median area; scutellum smooth, scutellar sulcus deep and scrobiculate, with six longitudinal carinae; mesopleuron largely smooth, striate near subalar furrow; posterior mesopleural sulcus distinct and scrobiculate; subalar groove scrobiculate; precoxal sulcus smooth; metapleuron smooth; propodeal areola distinctly delimited by carinae and with crossing transversal rugae. Legs: fore tibia with a row of 15 spines; hind coxa striate dorsally, smooth ventrally, with a distinct basoventral tooth. Wings: fore wing 2.8 times longer than wide; pterostigma 0.6 times the length of vein R; vein r 1.5 times longer than vein 3RSa; vein 3RSa 0.8 times as long as vein r-m; vein m-cu distinctly antefurcal, vein (Rs+M)b present, 0.2 times length of vein 2RS; hind wing with vein M+CU as long as vein 1M and twice length of vein r-m; vein m-cu slightly curved towards wing apex. Metasoma: first metasomal tergite short, about as long as its apical width, with median dorsal area delimited by two dorsolateral carinae, median area rugulose-coriaceous, lateral areas striate; second metasomal tergite striate medially, with coriaceous sculpture in between striations, with two more conspicuous slightly convergent furrows; third metasomal terigte smooth, turning coriaceous later ally, with a transverse scrobiculate furrow; remaining metasomal tergites smooth and polished; ovipositor 0.5 times length of metasoma, without distinct nodes.

Male. Smaller than female, body length 2.7-2.8 mm.

Variation. Female: Body length 4.1-4.6 mm; eyes 1.3-1.4 times higher than wide (lateral view); malar space 0.1-0.2 times eye height (lateral view); ocello-ocular distance 0.7-0.8 times diameter of lateral ocellus; antennae with 28-29 flagellomeres; fore wing length 2.8-2.9 times its maximum width; length of pterostigma 2.6-2.8 times its maximum width; hind wing vein M+CU 1.0-1.1 times longer than vein 1M.

Holotype. IB-UNAM CNIN. Female. Mexico, Jalisco, Estación Biológica Chamela, cerca laboratorio, 19.49, -105.04, 5.v.2011, 99-122 msnm, light trap, selva baja caducifolia, Cham-084, Zaldívar, Zaragoza, Ibarra col. DNA voucher no. CNIN795, GenBank accession nos JQ268745, JQ268749 (IB-UNAM CNIN).

Paratypes. IB-UNAM CNIN, MACN, UWIM. Three specimens. One female, Mexico, Jalisco, Estación Biológica Chamela, camino Búho, 19.49/19.49 N, -105.04/-105.04 E, 25.ii.2010, 106 msnm, sweeping net, selva baja caducifolia, A. Zaldívar col., DNA voucher no. ASDOR761, GenBank accession no. HQ200977; two males, Mexico, Jalisco, Estación Biológica Chamela, cerca laboratorio, 19.49 N, -105.04 E, 20.ii.2011, 99-122 msnm, light trap, selva baja caducifolia, A. Zaldívar, col., DNA voucher nos A SDOR 551 (Cham-500), ASDOR555 (Cham-504), GenBank accession nos HQ200979, HQ200967 (IB-UNAM CNIN).


The specific epithet derives from the greek words makros and ophthalmos, in reference to the very large compound eyes of this species.