Clubiona filifera Dankittipakul, 2008

Dankittipakul, Pakawin, Tavano, Maria, Chotwong, Wimolwan & Singtripop, Tippawan, 2012, New synonym and descriptions of two new species of the spider genus Clubiona Latreille, 1804 from Thailand (Araneae, Clubionidae), Zootaxa 3532, pp. 51-63 : 57-58

Clubiona filifera Dankittipakul, 2008

( Figs 18–24 )

Clubiona filifera Dankittipakul. Dankittipakul & Singtripop, 2008a: 41 , figs 20–21, 48–51 , description of male.

Type material. Holotype: 3, THAILAND: Nakhorn Ratchasima Province: Khao Yai National Park , Khao Yai , forests surrounding park headquarters, 800–1000 m, 15 September 2006, leg. P. Dankittipakul ( MHNG, examined) . New material: THAILAND, Nakhorn Ratchasima Province: 1Ƥ, Khao Yai National Park, sweeping in forests, 12 August 2010, leg. S. Pimpisalee ( MHNG) . Chantaburi Province: 1Ƥ, mandarin orchard, 1988, no collector ( EZTH) . Chachoengsao Province: 1Ƥ, orchards, 1985, no collector ( EZTH)

Extended diagnosis. Males of C. filifera are recognized by their long, ribbon-shaped embolus, which is coiled and occupies most of the apical extent of the palpal bulb ( Figs 20–22 ). Males are distinguished from those of the closely related C. cirrosa Ono, 1989 by the spiniform tegular apophysis, and by the digitiform RTA. Females can be recognized by the broad and shallow epigynal atrium ( Figs 18 , 23 ), and by the seemingly elongated insemination ducts encircling the atrium ( Figs 19 , 24 ).

Remarks. As expected, the recently discovered females have elongated insemination ducts of corresponding length related to the male embolus. The modified margin of epigynal atrium is accommodated by an elevated ridge, which seems to be responsible for guiding an elongated embolus towards the anteriorly located copulatory orifices during copulation.

Description. Female: Total length 6.8; prosoma 2.7 long, 2.0 wide; opisthosoma 4.1 long, 2.3 wide. Eye sizes and interdistances: AME 0.12, ALE 0.14, PME 0.13, PLE 0.14, AME–AME 0.10, AME–ALE 0.08, PME–PME 0.28, PME–PLE 0.16. Leg formula 4213. Leg measurements. Leg I 7.16 (1.90, 1.10, 1.92, 1.26, 0.98), leg II 7.72 (2.04, 1.12, 2.12, 1.48, 0.96), leg III 5.94 (1.82, 0.60, 1.26, 1.54, 0.72), leg IV 8.84 (2.22, 1.24, 1.98, 2.26, 0.74). Coloration and pattern. Prosoma strongly convex, in profile highest between PME and fovea. Carapace pale yellow, marked with narrow longitudinal band running between PME and fovea; pair of purple, irregular-shaped patches situated behind ocular region. Opisthosoma elongate-ovoid; dorsum yellowish brown, with dark broken bands running on lateral sides; venter pale. Spinnerets dark brown, distinctly darker on dorsal side.

Genitalia ( Figs 18–19 , 23–24 ). Epigynal atrium slightly depressed, trapezoidal, margin of atrium distinctly elevated, posterior margin poorly delimited. Copulatory orifices situated anteriorly, hidden underneath elevated marginal ridge. Insemination ducts descending posteriorly in mid-longitudinal axis, abruptly ascending anteriorly then curving ectad, aligned horizontally and terminally connected to triangular spermathecal heads. Fertilization ducts lanceolated.

Natural history. Clubiona filifera seems to prefer shady habitat; all specimens were collated in forests and orchards.

Distribution. Eastern Thailand.

Figures

FIGURES 11 – 19. Clubiona spp. 11 – 13. C. digitata sp. nov., holotype male. 14 – 16. C. aberrans sp. nov., holotype male. 17. C. digitata sp. nov., paratype female. 18 – 19. C. filifera Dankittipakul, female. 11 – 16. Male palp: ventral (11, 14), prolateral (12, 15), retrolateral (13, 16). 17. Right internal genitalia, dorsal. 18. Epigyne, ventral. 19. Internal genitalia, dorsal.

FIGURES 20 – 24. Clubiona filifera Dankittipakul, holotype male (20 – 22) and female (23 – 24). 20 – 22. Male palp: ventral (20), prolateral (22), retrolateral (21). 23. Epigyne, ventral. 24. Internal genitalia, dorsal.