Verrucidae Darwin, 1854

Buckeridge, John S., Jagt, John W. M. & Speijer, Robert P., 2008, Verruca punica, a new species of verrucomorph barnacle (Crustacea, Cirripedia, Thoracica) from the Lower Danian (Palaeocene) of Tunisia, Zootaxa 1844, pp. 37-46 : 39

Family Verrucidae Darwin, 1854

Diagnosis. Wall asymmetric, comprising four immovable dissimilar calcareous plates: carina, rostrum, scutum and tergum; second scutum and tergum together form a movable operculum; basis membranous or calcareous.

Remarks. In addition to the Verrucidae , the Verrucomorpha comprises two other families, the earliest of which, the Proverrucidae Newman, 1989 ( Newman and Hessler, 1989) , (Cenomanian-Upper Maastrichtian), is characterised by the presence of extra (lateral) plates in the shell wall, and the extant Neoverrucidae Newman in Newman & Hessler, 1989, which is a true ‘link’ with early non-verrucomorph barnacles, as it retains several basal whorls of imbricating plates, demonstrating phyletic proximity to a scalpelliform ancestor ( Newman, 2000; Buckeridge & Newman, 2006).