Rosapha bimaculata Wulp

Kovac, Damir & Rozkošný, Rudolf, 2012, A revision of the genus Rosapha Walker (Diptera: Stratiomyidae), Zootaxa 3333 (1), pp. 1-23 : 4-5

Rosapha bimaculata Wulp in de Meijere

( Figs 1–7 , 55–56 , 80 )

Rosapha bimaculata Wulp in de Meijere, 1904: 96

Diagnosis. Basal part of flagellum subconical, last flagellomere almost aristate but densely short pilose and thus appearing cylindrical, frontal tomentose spots absent, scutum and scutellum reddish yellow, wing with two separated brownish spots, legs yellow except for apex of hind femur and hind tibia, also fore tarsi and apices of other tarsi darkened.

Description. Male ( Figs 1–2, 4–7 , 55–56 ). Length: body 6.6–8.4 mm, wing 5.8–7.2 mm. Head ( Fig. 2 ) as in other species of genus, somewhat higher than long due to markedly prominent ocellar triangle in profile and about 1.5 times as broad as long in dorsal view. Vertex posterior to ocellar triangle virtually absent, upper frons narrow, barely longer than ocellar triangle. Lower frons transversely subtriangular, entirely shining black, without tomentose spots. Basal antennal segments and basal part of flagellum reddish yellow, apical style black ( Fig. 1 ). Basal part of flagellum subconical, last flagellomere about 1.9–2.1 as long as rest of antenna. Whitish tomentose stripes along inner margin of eyes with white hairs, in upper part almost as broad as scape. Central area of face with erect black hairs, lower part of head and postgena with erect white hairs. Labellum brownish at least in distal half, palpus yellow but greater part of its distal segment black.

Thorax reddish yellow including scutellum and upper half of pleura ( Figs 55–56 ). Scutum with a keel-shaped blackish median spot at anterior margin, also lower half of pleura black. Postpronotal callus and narrow subnotopleural stripe pale yellow, scutellum as in Fig. 4 . Wings with two brown, well separated areas: stigma and subapical band from apices of R4 and R5 to M1 or slightly beyond it. Legs yellow, but apex of hind femur and mid and hind tibiae black. Fore tarsus and last 3 apical tarsomeres of mid leg and 4 apical tarsomeres of hind leg brown, mid and hind basitarsus contrastingly whitish yellow.

Abdomen dark brown, often more reddish basally, and with brownish band along tergites laterally. Dense appressed black pubescence developed dorsally and white pubescence ventrally, longer dark hairs visible along lateral margin. Male terminalia ( Figs 5–7 ): Proctiger stout, almost pentagonal, cerci relatively short, genital capsule with proximal prominence and subtriangular, apically rounded posterior medial process, gonocoxite with prominence at inner posterior angle, gonostylus with broad submedial spine. Aedeagal complex similar to R. habilis (cf. Fig. 24 ).

Female ( Fig. 3 ). Length: body 5.2–6.6 mm, wing 4.7–6.5 mm. Head dichoptic, about 1.6 times broader than long in dorsal view. Frons at narrowest point about 1.5 times as broad as ocellar triangle, frons above antennae shining black, postocular band (in dorsal view) narrower than scape is long. Basal part of flagellum more oval than in male, last flagellomere about 1.7 times as long as rest of antenna. Tomentose facial stripes as in male, labellum more extensively brown. Medial spot at anterior margin of scutum sometimes replaced by transverse dark stripe.

Variation. The darkening of the male pleura is sometimes extensively reduced, often only the ventral part of the katepisternum is dark and the wing spots may be slightly more extensive. Scutellar spines are often paler than the scutellum. The pale abdominal lateral band may be indistinct.

Material examined. Type material: The male holotype should be deposited in ZMAN ( Woodley 2001) where it was not found (recent communication by B. Brugge), but it was figured in the original description. It was described from " Java , West Preanger , Gunong Tji Salimar , Kannegieter leg."

Other material: India: Assam, 10 miles N of Tinsukia, dense jungle, 29. iii.1944, 1 3, D.E. Hardy, in BPBM. Meghalaya, Nongpoh Forest, 25–28. iv. 1980, 1 ♀, A. Freidbeg, in USNM. Laos: Louang Phrabang Prov., Ban Song Cha (5 km W), 1200 m, 20°34´N, 102°14´E, 24.iv.–16. v.1999, 1 ♀, V. Kubáň, in MMB. Malaysia: Pahang distr., 30 km NE Raub, Lata Lembik, 3°56´N, 101°38´E, 200–400 m, 22.iv.–15. v.2002, 1 ♀, E. Jendek & O. Šauša, in ZIB. Thailand: Chiangdao, 5.–11.iv.1958, T. C. Maa, 1 3 in BPBM, 3 3 in USNM; Trang Province, Khaophappha, Khaochang, 200–400 m, 11. i.1964, 1 ♀, G. H. Samuelson, in USNM; Mae Hong Son, District Pangmapha, near Pai Viewpoint [Kiew Lom], ca. 1300 m, sweeping along stream, 12. iv.2010, 1 3, D. Kovac, in FSMU + 3 3, D. Kovac, in SMF; Mae Hong Son, Pangmapha, Pai Viewpoint [Kiew Lom], sweeping along stream, between 1400–1500 m, 28. iv.2010, 1 3, 1 ♀, D. Kovac, in SMF; Mae Hong Son, Pangmapha, Viewpoint [Kiew Lom] towards Pai, sweeping along stream, 22. iv.2010, 2 3, D. Kovac, in SMF; Mae Hong Son, Pangmapha, near Ban Nam Rin, Rudi valley, sweeping along right stream, 26. iv.2010, 2 3, D. Kovac, in SMF; Mae Hong Son, Pangmapha, Viewpoint [Kiew Lom] towards Pai, sweeping along stream, 1400–1500 m, 28. iv.2010, 1 3, 1 ♀, D. Kovac, in SMF; Mae Hong Son, Pangmapha, Viewpoint [Kiew Lom] towards Pai, sweeping along stream, c. 1650 m, 28. iv.2010, 1 3 (head missing), D. Kovac, in SMF; Mae Hong Son, Pangmapha, Pai Viewpoint [Kiew Lom], sweeping from bamboo leaves, 1621 m, 30. iv.2010, 1 3, D. Kovac, in SMF. Vietnam: Ninh Binh, Cuc Phuong N. P. near entrance, ca. 225 m, Malaise trap, 1.–15. v.2000, 1 ♀, Mai Phu Quy, in RMNH.

Distribution. Described from Indonesia (Java), our records extend the known distribution also to India, Malaysia, Thailand and Vietnam ( Fig. 80 ).

Figures

FIGURE 80. Distributional map of Rosapha bimaculata, R. brevispinosa sp. nov., R. obscurata, R. stigmatica sp. nov., and R. variegata. Some locality symbols represent several closely located collecting sites.

FIGURES 1 – 11. Diagnostic characters of Rosapha bimaculata and R. brevispinosa sp. nov. 1 – 7, R. bimaculata, male antenna (1), male and female head in frontal view (2 and 3), male scutellum (4), male terminalia: dorsal part (5), genital capsule (6) and gonostylus in posterolateral view (7). 8 – 11, R. brevispinosa sp. nov., male terminalia: dorsal part (8), genital capsule (9), gonostylus in posterolateral view (10), aedeagal complex (11). Scale bars: 1.0 mm (Figs 2 and 3), 0.5 mm (Figs 1 and 4), 0.3 mm (Figs 5 – 11).

FIGURES 55 – 62. Diagnostic characters of Rosapha bimaculata, R. brevispinosa sp. nov., R. flagellicornis, and R. flavipes sp. nov. 55 – 56, R. bimaculata, male, dorsal view (55) and lateral view (56). 57 – 59, R. brevispinosa sp. nov., female, dorsal view (57), lateral view (58) and frontal view (59). 60, R. flagellicornis, female, dorsal view. 61, R. flavipes sp. nov., male, dorsolateral view. 62, R. flavistigmatica sp. nov., male, lateral view.

FIGURES 22 – 32. Diagnostic characters of Rosapha habilis and R. handschini comb. nov. 22 – 26, 30, R. habilis, male terminalia: dorsal part (22), genital capsule (23), aedeagal complex (24), female terminalia in dorsal view (25), genital furca (26), gonostylus in posterolateral view (30). 27 – 29, 31 – 32, R. handschini comb. nov., male terminalia: dorsal part (27), genital capsule (28), aedeagal complex (29), gonostylus in posterolateral view (31), male antenna (32). Scale bars: 0.3 mm (Figs 22 – 26, 30), 0.2 mm (Figs 27 – 29, 31), 0.5 mm (Fig. 32).