Zoosphaerium libidinosum (de Saussure & Zehntner, 1897)

Wesener, Thomas & Wägele, Johann-Wolfgang, 2008, The giant pill-millipedes of Madagascar: revision of the genus Zoosphaerium (Myriapoda, Diplopoda, Sphaerotheriida), Zoosystema 30 (1), pp. 5-85 : 21

publication ID

https://doi.org/ 10.5281/zenodo.5391729

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scientific name

Zoosphaerium libidinosum


Zoosphaerium libidinosum View in CoL species-group

SPECIES INCLUDED. — Zoosphaerium libidinosum (de

Saussure & Zehntner, 1897); Z. arborealis Wesener & Sierwald , 200; Z. lambertoni ( Brolemann, 1922) ; Z. priapus (de Saussure & Zehntner, 1897) ; Z. anomalum (de Saussure & Zehntner, 1902) .

DIAGNOSIS. — These Malagasy giant pill-millipede species share a number of characters, identifying them as a species-group. Species of this group can be identified by a striking similarity in the shape of posterior telopods ( Figs 9K View FIG ; 14K View FIG ; 17I View FIG ). The third joint carries always more than 1 large, sclerotized, crenulated teeth on its backside closely to the tip. The hollowed-out margin juxtaposed to the second joint carries 2 large, membranous lobes and some shorter sclerotized spines ( Figs 9K View FIG ; 14L View FIG ; 17J View FIG ). To this species-group belong the only Zoosphaerium taxa, which (still?) possess a well-rounded operculum ( Fig. 9C, D View FIG ). Inside the group the species are clearly distinguishable by numerous characters, like the shape of anterior pair of telopods, the number of sensory cones on antenna, sensory cones on gnathochilarium, special shape of the female vulva operculum, body size, special bristle patterns on the collum and different structures on the surface of tergites.


A phylogenetic analysis will be necessary to confirm if the libidinosum -group represents a monophyletic group to be kept separately from the other Malagasy sphaerotheriids.

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