Cyphochilus feae Brenske, 1903

Sabatinelli, Guido, 2020, Taxonomic notes on the genus Cyphochilus Waterhouse, 1867 (Coleoptera, Scarabaeoidea, Melolonthinae) with description of 10 new species, Revue suisse de Zoologie 127 (1), pp. 157-181 : 170

Cyphochilus feae Brenske, 1903

Figs 10 , 65-67 , 83

Cyphochilus feae Brenske, 1903: 380

Type material examined: Holotype: ZMHB; 1 ♂; MYANMAR: // Feae 1092 / Type Brsk ( H) // Carin Cheba / 900-1100 m / L. Fea V XII-88 ( P) // coll. Brenske ( P) //. ( Figs 10 , 83 ).

Addition material examined: 11 specimens.

MYANMAR: ZMHB; 1 ♂; same data as holotype . – MNHN; 1 ♂; same data as holotype . – MSNG; 4 ♂; same data as holotype . – MHNG; 2 ♂; same data as holotype . – PCMN; 3 ♂; SE Myanmar, Dawna ; IX.1990.

Redescription of the holotype male: Size – BL: 14.5 mm; BW: 6 mm; BWX: 7.3 mm, situated at midpoint of elytra. Color. Integument dark shiny brown; dorsal surface with scattered yellow scales not aggregated along lateral part of the body. Head – CW/L: 3.6; anterolateral edges of clypeus rounded, lateral edges of clypeus curved and divergent posteriad; frons large, making eyes relatively small, not prominent (F/O: 3.9); antennal club longer than antennomeres 2-7 (A2-7/CL: 0.89). Pronotum – Transverse (PnW/L: 2.3), strongly convex; apical angles and lateral margins flattened, apical angles right, not protuberant; basal lateral margins straight, basal angles obtuse and largely rounded. Elytra – Without any visible striae. Thoracic sterna – Surface with short, white pubescence and scales; mesosternum slightly developed anteriorly (MstL: 0.24 mm) however, visible in lateral view between the mesocoxae. Abdomen – Sternites with sparse scales; pygidium convex with apical margin narrowly reflected. Legs – Protibia tridentate with the basal tooth well developed; slender apical tooth longer than the middle one. Aedeagus – Parameres strongly asymmetric with 5 appendices ( Figs 65-67 ).

Variability: BL: 7-18 mm. It is possible that the scattered scales on the dorsal part of the body in the specimens examined are a consequence of mechanical wear and that fresh specimens may have a denser covering of scales.

Females: Unknown.

Type locality: The type locality Carin Cheba corresponds nowadays to Karen Hills (750 m altitude) in Myanmar.

Distribution: Apart from Karen Hill, this species is known to me only from one other locality (Dawna Range) also situated in the Kayah State of eastern Myanmar.

Remarks: Cyphochilus feae , C. proximus and C. rohingyae have a similarly shaped parameres with five appendices and they are all known from Myanmar and the Malaysian Peninsula. They can be readily separated from each other mainly by the different conformation of the distal part of the right paramere ( Figs 53 , 65 , 71 ).

Figures

Figs 1-14. Original labels of type material. (1) Cyphochilus pygidialis Nonfried, 1893. (2) C. pygidialis Brenske [sic]. (3) C. septentrionalis Waterhouse, 1867. (4) C. obscurus Sharp, 1876. (5) C. proximus Sharp, 1876. (6) C. flavomarginatus Frey, 1971. (7) C. cylindricus Brenske, 1903. (8) Melolontha candida (Olivier, 1789). (9) Cyphochilus pygidialis v. angeri Nonfried, 1893. (10) C. feae Brenske, 1903. (11) C. vestitus Sharp, 1876. (12) C. tricolor Waterhouse, 1867. (13) C. testaceipes Fairmaire, 1902. (14) C. peninsularis Arrow, 1938.

Figs 59-70. Parameres in right, dorsal, and left view. (59-61) Cyphochilus zuercheri n. sp., holotype. (62-64) C. reichenbachi n. sp., holotype. (65-67) C. feae Brenske, 1903, holotype. (68-70) C. leducthoi n. sp., holotype. – Scale bars = 1 mm.

Figs 80-91. Habitus. (80) Cyphochilus candidus (Olivier, 1789), lectotype ♂ MNHN. (81) C. candidus (Olivier, 1789), Nepal: Godavari, ♂ MHNG, yellow scales aberration. (82) C. carinchebanus Brenske, 1903, Myanmar: Carin Cheba, ♂ ZMHB. (83) C. feae Brenske, 1903, holotype ZMHB. (84) C. gandhii n. sp., holotype MHNG. (85) C. leducthoi n. sp., holotype MHNG.(86) C. septentrionalis Waterhouse, 1867, lectotype BMNH. (87) C. obscurus Sharp, 1876, holotype MNHN. (88) C. sansuukyii n. sp., paratype ♀ MHNG. (89) C. peninsularis Arrow, 1938, paralectotype ♂ MHNG. (90) C. rohingyae n. sp., holotype MHNG. (91) C. rohingyae n. sp., paratype ♂ MHNG, Malaysia: Perak, bicolor aberration. – Scale bars = 5 mm.

Figs 47-58. Parameres in right, dorsal, and left view. (47-49) Cyphochilus carinchebanus Brenske, 1903. (50-52) C. obscurus Sharp, 1876, holotype. (53-55) C. proximus Sharp, 1876, holotype. (56-58) C. tricolor Waterhouse, 1867, lectotype. – Scale bars = 1 mm.

Figs 71-79. Parameres in right, dorsal, and left view. (71-73) Cyphochilus rohingyae n. sp., holotype. (74-76) C. hmong n. sp., holotype. (77-79) C. orbachi n. sp., holotype. – Scale bars = 1 mm.