Stenus, Puthz, 2013

Puthz, V., 2013, Übersicht über die orientalischen Arten der Gattung Stenus LATREILLE 1797 (Coleoptera, Staphylinidae) 330. Beitrag zur Kenntnis der Steninen, Linzer biologische Beiträge 45 (2), pp. 1279-1470 : 1284-1293

publication ID

https://doi.org/ 10.5281/zenodo.5300922

DOI

https://doi.org/10.5281/zenodo.5307845

persistent identifier

https://treatment.plazi.org/id/A92887EE-FFA8-7020-FEE0-FCC55E507302

treatment provided by

Marcus

scientific name

Stenus
status

 

Key to the groups of the Oriental species of the genus Stenus View in CoL

1 Abdominal segments 4-6 margined (margin can be thread-like).........................................2

- Abdominal segments 4-6 immargined.................................................................................3

2 Tarsi simple (segment 4 not bilobed) .......................................................................... key A

- Tarsi bilobed (segment 4 bilobed)............................................................................... key B

3 Tarsi simple....................................................................................... S. perplexus View in CoL -group pp.

- Tarsi bilobed ............................................................................................................... key C

A. Key to the Oriental species of the genus Stenus LATREILLE with distinct paratergites and simple (not bilobed) tarsi exclusively brachypterous/apterous montane Chinese species

1 Base of abdominal tergites without short carinae................................................................2

- Base of abdominal tergites with short carinae...................................................................45

2 Sternum 9 with an acute and prominent apicolateral tooth (figs 18-20)..............................3

- Sternum 9 serrate apicolaterally (this character may be difficult to find), posterior margin of the sternite finely serrate (figs 21-26). Median lobe with ( S. pusillus View in CoL -group) or without expulsion claps or expulsion hooks (figs 85, 86) ( S. seminiger View in CoL -group, S. comma-group)...................................................................................................................33

3 Abdomen with conspicuously long and erect setae (figs 6, 7).: Aedeagus (fig. 14, PUTHZ 2009c). China (Taiwan). (S. cirrus- group p.p.)....................... S. cirrivarians PUTHZ View in CoL

- Abdomen not with conspicuously long and erect setae.......................................................4

4 Smaller species: FB <2.6 mm. Posterior margin of the sternite ± smooth. Median lobe with expulsion clasp or expulsion hooks.............................................................................5 - Larger species (FB>2.5mm). Posterior margin of the sternite distinctly or very finely serrate (figs 22, 23, 25, 26). Median lobe without expulsion clasps or expulsion hooks ( S. seminiger View in CoL -group p.p.....................................................................................................26

5 Small metallic species.: Apical portion of median lobe setaceous.: Spermatheca sclerotized ( S. viridescens View in CoL -group).......................................................................................6

- Blackish species of various length.: Apical portion of median lobe not setaceous.: Spermatheca sclerotized or not sclerotized....................................................................7

6 Larger: 3.0-3.5mm.: Aedeagus (fig. 70). N India....................... S. viridescens CAMERON View in CoL

- Smaller: 2.3-2.8mm.: Aedeagus (fig. 71).: Spermatheca (fig. 77). (N India) ............... ....................................................................................................... S. almorensis CAMERON View in CoL

7 Brachypterous species, FB 1.6mm.: Spermatheca sclerotized........................................ .......................................................................... S. alpicola View in CoL -group and S. brahmanus View in CoL -group S. dodabettamontis PUTHZ.: Unknown.: Valvifer ( Abb. 27 View Abb ). S India; S. morosus CAMERON. View in CoL : Aedeagus resembling fig. 72). India, Pakistan; sikh PUTHZ.: Aedeagus (figs 4, 5, PUTHZ 1985b). India, Pakistan; S. brahmanus View in CoL -group (see below)

- Macropterous species ..........................................................................................................8

8 Species with long conspicuous recumbent pubescence, especially distinct on abdomen.: Aedeagus without a long flagellum or a long median tube.: Spermatheca not sclerotized. (mendicus -group =) ................................................................ S. incanus View in CoL -group S. eurous PUTHZ View in CoL (fig. 18, l. c.).: Aedeagus (figs 33-35, l. c.). China, Vietnam, Laos; S. fukiensis L. BENICK. View in CoL : Aedeagus (fig. 31, l. c.). China; S. insulindicola PUTHZ. View in CoL : Aedeagus (fig. 82, l. c.). Indonesia: Sumatra; S. insulindicus ROUGEMONT. View in CoL : Aedeagus (fig. 83, l. c.). Indonesia: Bali; S. koreanus PUTHZ. View in CoL : Aedeagus (fig. 19, l. c). Korea; S. lanicutis PUTHZ. View in CoL : Aedeagus (figs. 22, 32, 36, l.c.). China; S. lanosivestis PUTHZ. View in CoL : Aedoeagus (fig. 7, PUTHZ 2012c). Myanmar; S. lanosus PUTHZ. View in CoL : Aedeagus (figs. 78-81, l. c.). China; S. puberulus SHARP. View in CoL : Aedeagus (figs 23-28, PUTHZ 2012a). Japan; S. pubiformis PUTHZ. View in CoL : Aedeagus (figs. 20, 29, 30, l.c.). China, Korea, Russia.

- Species without conspicuous pubescence.: Spermatheca sclerotized..............................9

9: Aedeagus with a long flagellum or a long median tube.: Spermathecal duct slender (fig. 83).................................................................................................................10

-: Aedoeagus without a long flagellum (figs 72, 82).: Spermatheca sclerotized, spermathecal duct stout. 3.8-4.4mm (FB 2.0-2.4mm). Pakistan, Indien, Nepal, Bhutan....... .......................................................................................................... S. musicola CAMERON View in CoL

10 Smaller species: FB <1.8mm.: Median lobe simple posteriorly S. javanus View in CoL -group........11

- Larger species: FB>1.9mm.: Median lobe with the internal tube/flagellum projecting posteriorly (figs 79-81) S. flagellifer View in CoL -group.....................................................21

11 Larger: FB 1.6mm...........................................................................................................12

- Smaller: FB <1.6mm.........................................................................................................13

12 Punctation of elytra and pronotum less coarse, mostly distinctly delimited. Abdominal punctation denser, interstices on posterior portion of tergites at most as large as punctures.: Apical portion of median lobe acute, much longer than the parameres (fig. 78). 3.3-4.2mm (FB 1.8mm). Myanmar.......................................... S. collium FAUVEL View in CoL

- Punctation of elytra and pronotum coarser, slightly rugose. Abdomen less densely punctate, interstices on posterior portion of tergites larger than punctures.: Aedeagus (fig. 2, PUTHZ 1969), median lobe much shorter than parameres. 3.3-4.2mm (FB 1.8mm). Myanmar, Thailand, Laos........................................................... S. comes FAUVEL View in CoL

13 Elytra with a reddish yellow spot......................................................................................14

- Elytra blackish. Several very similar species which should be identified by their sexual characters..........................................................................................................................15 14 Elytral spots larger, about as long as antennal segments 3-5.: Aedeagus (fig. 1, ROUGEMONT 1987), apical portion of median lobe about half as long as in S. pederseni View in CoL , with narrow ventral carinae. 2.7-3.3mm (FB 1.5mm). Thailand........................................... ................................................................................................... S. kamhaengi ROUGEMONT View in CoL

- Elytral spots smaller, about as long as antennal segment 3. Head broader (HW: EW 1.00).: Aedeagus (fig. 73).: Spermatheca (fig. 83). 2.7-3.3mm (FB 1.5-1.6mm). Laos....................................................................................................... S. pederseni View in CoL nov.sp.

15 Smaller: 2.0-2.7mm (FB 1.3-1.4mm).: Aedeagus (figs 12, 15, PUTHZ 2012a), apical portion.: Spermatheca (fig. 16, l. c.). China (Taiwan), Japan (Ryukyu Islands), Malaysia (Sabah), Laos........................................................................ S. riukiuensis PUTHZ View in CoL

- Larger: 2.4-3.4mm. Aedeagus different............................................................................16

16 Head broader (HW: EW  1.0).........................................................................................17

- Head less broad (HW: EW>0.95).....................................................................................20

17 Punctation of elytra coarser, diameter of punctures larger than diameter of antennal segment 1.: Median lobe closely resembling that of S. riukiuensis View in CoL , but the apical portion shorter, parameres about as in S. sannio View in CoL ( PUTHZ 2012a, fig. 15). 2.4-3.0mm (FB 1.4-1.5mm). Malaysia (Pahang, Sabah). .................................. S. spongifera CAMERON

- Punctation of elytra less coarse, diameter of punctures less large than diameter of antennal segment 1............................................................................................................18

18. Aedeagus (fig. 73a). 2.3-3.0mm (FB 1.4-1.6mm). Indonesia (Java)................................ ......................................................................................................... S. javanus BERNHAUER View in CoL

-: Aedeagus otherwise.....................................................................................................19

19: Aedeagus (fig. 74). 3.0mm (FB 1.5mm). N India.............................. S. lescheni View in CoL nov.sp.

-: Aedoeagus (fig. 75). 2.4-3.1mm (FB 1.4-1.6mm). Indonesia (Lombok, Java)................ ................................................................................................................. S. sannifer PUTHZ View in CoL

20 Abdominal punctation denser, interstices on tergite 7 at most as large as half diameter of punctures.: Median lobe (fig. 6, PUTHZ 1991) closely resembling that of S. javanus View in CoL .: Spermatheca (fig. 7 l. c.). 3.4mm (FB 1.5-1.6mm). N India, Nepal................... ................................................................................................................ S. sannator PUTHZ View in CoL

- Abdominal punctation slightly less dense, interstices on tergite 7 occasionally as large as half diameter of punctures or slightly larger.: Aedeagus (fig. 6, PUTHZ 1980a). 2.7- 3,3mm (FB 1.5-1.6mm). N India, Nepal, China (Yunnan) ................... S. sannio PUTHZ View in CoL

21 Legs dark...........................................................................................................................22

- Legs reddish yellow or reddish brown..............................................................................23

22 Head narrower (HW: EW<0.95). Elytra shorter (EW: EL 1.00).: Anterior portion of median lobe triangular (fig. 1, PUTHZ 1976a). Indonesia (Java) ........ S. flagellifer PUTHZ View in CoL

- Head slightly broader (HW: EW>0.95). Elytra longer (EW: EL <0.95).: Aedeagus (fig. 89), median lobe with a long, strongly sclerotized tube. 4.2mm (FB 2.2mm). Indonesia (Sulawesi) ........................................................................ S. sulawesicola View in CoL nov.sp.

23 Abdominal punctation dense or moderately dense............................................................24

- Abdominal punctation very sparse.: Aedeagus (fig. 2, PUTHZ 1980b). 4.0-4.5mm (FB 2.2mm). Philippines (Luzon, Negros, Imugan). .......................... S. aeratus L. BENICK View in CoL

24 Abdominal tergites 3-6 with faint groundsculpture. Elytra about as long as broad. On average smaller.................................................................................................................25 Abdominal tergites 3-6 brilliant, without groundsculpture. Elytra longer than broad.: Aedeagus (fig. 88). 3.8-4.6mm (FB 2.5-2.6mm). Indonesia (Sulawesi, Maluku) .................. ............................................................................................................ S. quaesitus View in CoL nov.sp.

25: Aedeagus (figs 80, 81). 3.7-4.3mm (FB 1.9-2.0mm). Malaysia (Pahang), Indonesia Sumatra) ........................................................................................... S. pastoralis CAMERON View in CoL

-: Aedeagus (fig. 79). 3.3-4.1mm (FB 2.0-2.2mm). Malaysia (Sabah, Sarawak) ................ ..................................................................................................................... S. heissi PUTHZ View in CoL

26 Elytral sculpture strongly vorticose...................................................................................27 - Elytra sculpture not vorticose, punctures mostly distinctly delimited...............................29

27 Abdominal punctation coarser, punctures in tergite 7 as large as basal cross section of antennal segment 3.: Aedoeagus (fig. 5, PUTHZ 1975). 4.8-6.7mm (FB 2.7-3.3mm). N India, Nepal, Bhutan................................................................... S. sikkimensis CAMERON View in CoL

- Abdominal punctation less coarse, punctures on tergite 7 distinctly smaller than basal cross section of antennal segment 3..................................................................................28

28 Elytra subquadrate, broader (HW: EW <0.85).: Aedoeagus (fig. 85). 4.5-6.5mm (FB 2.7-3.2mm). N India, Nepal, China (Yunnan)................................... S. tortuosus CAMERON View in CoL

- Elytra subtrapezoid, less broad (HW: EW>0,90).: Aedoeagus (fig. 86). 5.0-6.0mm (FB 2.5-2.8mm). Nepal................................................................... S. sulcatipennis View in CoL nov.sp.

29 Punctation of frons less dense, interstices distinctly larger than half diameter of punctures. Aedoeagus (fig. 7, PUTHZ 1980b). 4.3-6.5mm (FB 2.6-2.8mm). Philippines (Luzon)............................................................................................. S. concavus L. BENICK View in CoL

- Punctation of frons very dense, interstices smaller than half diameter of punctures.........30

30 Paratergites less broad, those of tergite 4 about as broad as antennal segment 1..............31

- Paratergites broader, those of tergite 4 distinctly broader than antennal segment 1..........32

31 Smaller.: Aedoeagus (fig. 38, PUTHZ 2008b). 4.2-5.5mm (FB 2.5-2.6mm). China (Yunnan, Sichuan, Hunan) ................................................................... S. fraterculus PUTHZ View in CoL

- Larger.: Aedoeagus (fig. 39, PUTHZ 2008b). 4.8-6.2mm (FB 2.8-3.0mm). China (Taiwan) ................................................................................................ S. yanoianus PUTHZ View in CoL

32 Elytra broader.: Aedoeagus (fig. 44, PUTHZ 2008b). 4.8-6.2mm (FB 2.7-2.8mm). China (Sichuan)...................................................................................... S. atrovestis PUTHZ View in CoL

- Elytra less broad.: Aedoeagus (fig. 30, TANG & LI 2013). 67mm (FB 3.3mm). China (Ningxia) ...................................................................................... S. biwenxuani TANG & LI View in CoL

33 Larger species: FB>2.5mm..............................................................................................34

- Smaller species: FB <2.5mm............................................................................................36

34 Segment 1 of metatarsi shorter than the 3 following segments combined.........................35

- Segment 1 of metatarsi longer than the 3 following segments combined.........................37

35 Legs reddish brown. Abdomen finely punctate, punctures smaller than basal cross section of antennal segment 3.: Aedeagus (fig. 4A, ROUGEMONT 1983b). 4.8-5.5mm (FB 2.7-3.0mm). Thailand, Nepal, N India................... .............. S. lomholdti ROUGEMONT View in CoL

- Legs blackish. Abdomen coarsely punctate, punctures on tergite 7 as large as basal cross section of antennal segment 3.: Aedeagus (fig. 36, PUTHZ 2008b). 5.0-5.5mm (FB 2.5-2.7mm). China (Yunnan) ..................................................... S. tenebricosus PUTHZ View in CoL

36 Larger, with dark-blue metallic tint. Abdominal punctation very coarse, punctures on tergite 7 about as large as basal cross section of antennal segment 3.: Aedeagus (fig. 32, PUTHZ 1980b). 5.0-6.0mm (FB 3.0-3.2mm). China (Taiwan) ......................................... ................................................................................................... S. sauterianus BERNHAUER View in CoL

- Smaller, without blue metallic tint: Abdominal punctation finer, punctures on tergite 7 distinctly smaller than basal cross section of antennal segment 3.: Aedeagus (fig. 33, PUTHZ 2008b). 4.5-5.5mm (FB 2.4-2.7mm). Japan............................ S. kobensis CAMERON View in CoL

37 Smaller species: FB 1.8mm.............................................................................................38

- Larger species: FB>2.0mm. Aedeagus without an expulsion clasp.................................39

38 Smaller species: FB 1.5mm. Aedeagus with an expulsion clasp ( S. pusillus View in CoL -group: palaearctic) S. escensus PUTHZ. View in CoL : Aedeagus (fig. 9, PUTHZ 1970b). N India, Nepal; S. kishimotoianus PUTHZ. View in CoL : Unknown. China (Sichuan); S. pernanus PUTHZ. View in CoL : Aedeagus (fig. 6, PUTHZ 2006). China (Shaanxi); S. pusillulus PUTHZ. View in CoL : Aedeagus (fig. 1, PUTHZ 1986b). Pakistan; S. simlaensis CAMERON. View in CoL : Aedeagus (fig. 6, PUTHZ 1970b). N India; S. subescensus PUTHZ. View in CoL : Aedeagus (fig. 14, PUTHZ 1976b). Nepal;

S. tumidicollis PUTHZ. View in CoL Nepal; S. turgidicollis PUTHZ. View in CoL : Aedeagus (fig. 3, PUTHZ 1986b). Pakistan.

- Larger species: FB>1.5mm. Aedeagus expulsion hooks (fig. 7, PUTHZ 2002b). 2.6- 3.3mm (FB 1.6-1.7mm). Vietnam................ S. gibbicollis View in CoL -group: palaearctic) S. lac PUTHZ View in CoL

39 Elytra each with a reddish-yellow spot.............................................................................40

- Elytra without reddish spots..............................................................................................44

40 First segment of metatarsi distinctly longer than the three following segments combined...........................................................................................................................41

- First segment of metatarsi shorter or nearly as long the three following segments combined...........................................................................................................................42

41 Elytral spot very large, broader than half the breadth of elytron.: Aedeagus (fig. 34, PUTHZ 2008b). 4.5-6.0mm (FB 2.5-2.9mm). China (also Hainan) ........................................ .................................................................................................. S. huabeiensis ROUGEMONT View in CoL

- Elytral spot smaller than half the breadth of elytron.: Aedeagus (fig. 35, l. c.). 4.0- 5.0mm (FB 2.4-2.5mm). China (Sichuan)............................................ S. falsiloquax PUTHZ View in CoL

42 Lateral portions of frons sparsely punctate, interstices at least as wide as diameter of punctures...................................( S. comma View in CoL -group)...........................................................43

- Lateral portions of frons densely punctate, interstices smaller than diameter of punctures.: Aedeagus (fig. 12, Puthz 2008b). China (Yunnan, Sichuan).......................... .................................. ............................................................................ S. bioculatus PUTHZ View in CoL

43 Larger: 4.3-5.5mm. Elytral punctation very dense, punctures distinctly delimited, interstices shining.: Aedeagus (fig. 11, l. c.), apical setae of parameres very long 4.0-5.0mm. China (Fujian, Yunnan, Jiangsu, Guizhou, Jilin), Japan....... S. tenuipes SHARP View in CoL

- Smaller: 3.4-4.2mm. Elytral punctation extremely dense, rough, punctures less distinctly delimited, interstices totally dull (general impression: structure like charcoal).: Aedeagus (fig. 13, l. c.), apical setae of parameers short. China (Yunnan, Sichuan) ................................................................................................................................. ............................................................................................................. S. scabratus PUTHZ View in CoL

44 Abdomen nearly cylindrical, paratergites of tergite 4 narrower than antennal segment 2, directed ventrad.: Aedeagus (fig. 37, l. c.). 4.2-5.1mm (FB 2.4mm). China (Taiwan) ................................................................................................................................. ............................................................................................................. S. insulanus PUTHZ View in CoL

- Abdomen broadly-elliptical, paratergites broader, paratergites 4 as broad or broader thanantennal segment 2, directed dorsad or horizontally ...................................................... ................................( S. seminiger View in CoL -group, S. comma View in CoL -group p. p.: see PUTHZ 1980b, 2008b) S. confluens CAMERON. View in CoL : Aedeagus (fig. 11, PUTHZ 1980b). N India; S. erythrocnemus EPPELSHEIM. View in CoL : Aedeagus ( Abb. 2 View Abb , PUTHZ 1970d), Aserbeidschan, Armenia, Turkey, Iran, Tadzhikistan, Afghanistan, Pakistan; S. hirthei PUTHZ. View in CoL : Aedeagus (fig. 2, PUTHZ 2003b). Nepal; S. holzschuhi PUTHZ. View in CoL : Aedeagus (fig. 1, PUTHZ 1988b). Nepal; S. imitatus PUTHZ. View in CoL : Aedeagus (fig. 10, PUTHZ 1980b). N India; S. insignatus PUTHZ. View in CoL : Aedeagus (figs 16, 17, PUTHZ 20008). Vietnam, China (Hainan); S. insulanus PUTHZ. View in CoL : Aedeagus (fig. 1, PUTHZ 1971c). China (Taiwan); S. jaegeri PUTHZ. View in CoL : Aedeagus (fig. 3b, PUTHZ 2003b). Nepal; S. kurilensis PUTHZ. View in CoL : Unknown. Kuriles; S. nitidiceps PUTHZ. View in CoL : Aedeagus (fig. 15, PUTHZ 1980b). N India, Pakistan, Nepal; S. plumbativestis PUTHZ. View in CoL : Aedeagus (figs 22, 23, PUTHZ 2010). China (Yunnan); S. plumbatus PUTHZ. View in CoL : Aedeagus (fig. 14, PUTHZ 1980b). N India; S. plumbarius PUTHZ. View in CoL : Aedeagus (fig. 42, PUTHZ 2008b). China (Yunnan); S. plumbeus CAMERON. View in CoL : Aedeagus (fig. 1, PUTHZ 1968b; figs 12, 13, PUTHZ 1980b). Afghanistan, Pakistan, India, Nepal, Bhutan, China (Sichuan, Yunnan); S. plumbivestis PUTHZ. View in CoL : Aedeagus (fig. 41, PUTHZ 2008b). China (Taiwan); S. seminiger CHAMPION. View in CoL : Aedeagus (about as in fig. 10, PUTHZ 1980b). N India, Nepal, Bhutan.

45 Base of anterior tergites with 3 carinae.............................................................................46

- Base of anterior tergites with 4 carinae. 2.4-3.5mm (FB 1.4-1.7mm). India, Nepal, Myanmar, Thailand, Laos, Vietnam, China (Yunnan), Indonesia (Timor), Philippines....... .......................................................................................... S. melanarius View in CoL annamita FAUVEL

46 Tergite 10 with a spot of dense and shiny pubescence. Mostly smaller species (S. circularis- group [= S. rugicollis View in CoL -group]), key see PUTHZ 2003b S. adebratti View in CoL nov.sp..: Aedeagus (fig. 92). Malaysia (Sabah); S. adjunctus PUTHZ. View in CoL : Aedeagus (figs 9, 10, PUTHZ 1991). Nepal; S. baliensis ROUGEMONT. View in CoL : Aedeagus (fig. 1, ROUGEMONT 1984). Indonesia (Bali); S. beesoni CAMERON. View in CoL : Aedeagus (figs 14, 15, PUTHZ 1968b). N India, Myanmar, Thailand; S. boholensis View in CoL nov.sp..: Aedeagus (fig. 94). Philippines (Bohol); S. brookeianus ROUGEMONT. View in CoL : Unknown. Brunei; S. calcarifer PUTHZ. View in CoL : Aedeagus (figs 12, 13, PUTHZ 1991). N India; S. carinatus CAMERON. View in CoL : Aedeagus (fig. 1, PUTHZ 1971a). Sri Lanka; S. chatterjeei CAMERON. View in CoL : Aedeagus (fig. 2, l. c.). N India; S. concavifrons PUTHZ. View in CoL : Aedeagus (figs 9, 10, PUTHZ 2003b). Indonesia (Java); S. diffidens CAMERON. View in CoL : Aedeagus (figs 9, 10, PUTHZ 1968b). N India, Nepal, Myanmar, Thailand, Laos; S. duplopunctatus BERNHAUER. View in CoL Philippines (Luzon, Mindanao), Indonesia (Sumatra, Java), Malaysia (Sabah); S. falsidicus PUTHZ. View in CoL : Aedeagus (fig. 11, PUTHZ 1991). N India; S. floresicus PUTHZ. View in CoL : Aedeagus (fig. 12, PUTHZ 2003b). Indonesia ( Flores); S. geiseri View in CoL nov.sp..: Aedeagus (fig. 95). Laos; S. helleri L. BENICK. View in CoL : Aedeagus (fig. 13, PUTHZ 1968b). Indonesia (Sulawesi); S. helvitarsis PUTHZ. View in CoL : Unknown. Philippines (Mindoro) S. indicus PUTHZ. View in CoL : Aedeagus (figs 19, 20, PUTHZ 1968b). Bangladesh ("Bengala"),? China (Yunnan); S. indignandus PUTHZ. View in CoL : Aedeagus (fig. 7, PUTHZ 2003b). Indonesia (Sumatra); S. infaustus View in CoL nov.sp.: Aedeagus (fig. 96). Laos; S. jaechi PUTHZ. View in CoL : Aedeagus (fig. 8, l. c.). Malaysia ( Sarawak); S. louwerensi CAMERON. View in CoL : Aedeagus (figs 16, 17, PUTHZ 1968b). Indonesia (Java, Sulawesi),? Philippines (Luzon), Thailand, Laos, Vietnam; S. megacephalus CAMERON. View in CoL : Aedeagus (fig. 7A, ROUGEMONT 1981b). Malaysia, Indonesia (Sumatra, Java), Myanmar, Thailand, Laos, Vietnam, China (Yunnan); S. molestus PUTHZ. View in CoL : Aedeagus (figs 4, 5, PUTHZ 2003b). Nepal; S. mon ROUGEMONT. View in CoL : Aedeagus (fig. 10, ROUGEMONT 1981b). Thailand, Myanmar, Laos; S. notabilifrons View in CoL nov.sp.. Malaysia; S. rugicollis KRAATZ. View in CoL : Aedeagus (fig. 93). India, Myanmar, Thailand, Laos, Vietnam; S. skalei View in CoL nov.sp..: Aedeagus (figs 90, 91). Indonesia (Sulawesi); S. sumbaensis SCHEERPELTZ. View in CoL : Aedeagus (fig. 1, SCHEERPELTZ 1957). Indonesia (Sumba); S. timoricus PUTHZ. View in CoL : Aedeagus (fig. 3, PUTHZ 1966). Indonesia (Timor).

- Tergite 10 without a shiny spot of dense and shiny pubescence.......................................47

47 Sternum 9 apically serrate, smaller species..............................................( S. humilis View in CoL -group) No Oriental species, but several species in China and Japan

- Sternum 9 acute apicolaterally, larger species ( S. clavicornis View in CoL -group)...............................48 The following species of China, Korea and Japan, which belong to the palaearctic fauna are not included here: S. calliceps BERNHAUER View in CoL ( China, Japan), S. clavicornis (SCOPOLI) View in CoL (Holarctis, Japan), S. disterminus PUTHZ ( Japan) View in CoL , S. indagator EPPELSHEIM View in CoL ( Russia, China, Japan), S. kamtschaticus MOTSCHULSKY View in CoL ( China, Russia, Mongolia), S. lewisius SHARP ( Japan) View in CoL , S. lewisius View in CoL pseudoater BERNHAUER ( China, Korea), S. mongolicus EPPELSHEIM View in CoL ( China (Sichuan, Xizang, Qinghai, Gansu, Xinjiang), Kazakhstan, Turkmenistan, Tadzhikistan, Uzbekistan, Kyrgyzstan, Iran, Afghanistan, Pakistan, Ladakh, Kashmir, SE Russia), S. nishikawai NAOMI ( Japan) View in CoL , S. paradoxus Bernhauer View in CoL ( China, Russia), S. sauteri BERNHAUER View in CoL ( Japan, Korea, Russia), S. secretus BERNHAUER View in CoL ( Russia, China, Mongolia), S. ussuriensis RYVKIN View in CoL ( Russia, China), S. yamato NAOMI ( Japan) View in CoL (see PUTHZ 2013b)

48 Elytra each with a reddish yellow spot..............................................................................49

- Elytra without a reddish yellow spot.................................................................................50

49 Median portion of frons distinctly elevate, longitudinal furrows deep.: Apical portion of median lobe not broadened...............................................................................51

- Median portion of frons indistinctly elevated, longitudinal furrows shallow.: Apical portion of median lobe broadened (fig. 1, HROMÁDKA 1980). 4.5-5.3mm (FB 2.5- 2.6mm). N India, Pakistan, Afghanistan............................................ S. bengle HROMÁDKA View in CoL

50 Elytral spot in about middle of elytra (fig. 27, PUTHZ 2010).: Metafemora simple. Aedeagus (fig. 25, l. c.). 4.5-5.5mm (FB 2.3-2.5mm). N India, Pakistan, Nepal, China (Yunnan) ........................................................................................... S. kraatzi BERNHAUER View in CoL

- Elytral spot in posterior half of elytra (fig. 28, l. c.).: inner side of metafemora densely furnished with long setae. Aedeagus (fig. 24, l. c.). 4.8-6.0mm (FB 2.4- 2.6mm). China (Shaanxi, Sichuan) ................................................. S. lanuginosipes PUTHZ View in CoL

51 Paratergites broader, directed horizontally, paratergite 4 broader than antennal segment 1........................................................................................................................................52

- Paratergites less broad, directed ventrad, paratergite 4 at most as broad as antennal segment 1..........................................................................................................................53

52 Elytra with a small rosette of curved sulci medially.: Metafemur with a spine basally (fig. 29, l. c.). Aedeagus (fig. 26, l. c.). 3.8-4.5mm (FB 2.0mm). China (Sichuan, Yunnan)................................................................................................. S. spinulipes PUTHZ View in CoL

- Elytra without a rosette medially.: Metafemur simple. 3.5-4.2mm (FB 2.2mm). Aedeagus (fig. 2, PUTHZ 2006). Indonesia (Java)..................................... S. desertor PUTHZ View in CoL

53 Elytral sculpture strongly rugose/vorticose.......................................................................54

- Elytra very densely punctate, occasionally slightly rugose...............................................55

54 Head broader than elytra, paratergites indistinctly delimited, evanescent.: Aedeagus (fig.1, PUTHZ 2006). 4.0-5.2mm (FB 2.2mm). China (Taiwan) ............ S. pseudolus PUTHZ View in CoL

- Head narrower than elytra, paratergites narrow but distinctly delimited.: Aedeagus (fig. 84). 4.0-5.5mm (FB 2.7mm). China ( Taiwan, Hongkong, Fujian, Chongqing, Jiangsu), Vietnam, Japan............................................................... S. formosanus L. BENICK View in CoL

55 Abdominal punctation denser, tergite 7 matt, interstices much smaller than half diameter of punctures.: Median lobe slightly longer than parameres. 4.5-6.0mm (FB 2.1-2.6mm). Indien, Myanmar, Thailand, Vietnam................................. S. venator FAUVEL View in CoL

- Abdominal punctation less dense, tergite 7 shiny, interstices larger than half diameter of punctures.: Median lobe distinctly shorter than parameres. 4.0-5.0mm (FB 2.5mm). Japan, Korea, Russia, China (Fujian, Hainan, Hubei, Sichuan, Shaanxi, Beijing) ...................................................................................................... S. distans SHARP View in CoL

B. Key to the groups of Oriental species of the genus Stenus with margined abdomen and bilobed tarsi

1 Head with very large eyes, average breadth of frons at most half the width of head (figs 1, 2). Large species, FB> 2.0mm (habitus see also fig. 81, PUTHZ 1969). Paraglossae oval......................................................................................................................................2

- Head with smaller eyes (e. g. fig. 3), average distance between eyes broader than half the width of head. Species of various sizes. Paraglossae oval, coniform or sileniform.......4

2 Sternum 9 apicolaterally acute (figs 54, 55), frons deeply concace, median portion not or only slightly elevated......................................................................................................3 - Sternum 9 apicolaterally serrate (fig. 69), frons deeply concave, median portion strongly elevated .............................................................................................. S. feae View in CoL group

3: Legs simple, E: median lobe with a small triangular expulsion hook (e. g. figs 256- 260), parameres setaceous along whole internal side.: S (about as in figs 261, 262, 263, 273....................................................................................................... S. gestroi View in CoL -group

-: Legs with preapical/apical spurs, E: median lobe with a large expulsion clasp (e. g. figs 268-270, 275), parameres setaceous at apex.: S (about as in fig. 274)....................... ............................................................................................................. S. guttalis View in CoL -group pp.

4 Sternum 9 apicolaterally ± serrate (figs 35-38, 60, 64, 67, 68)...........................................5 (note: this characters sometimes is barely noticable: e. g. fig. 31)

- Sternum 9 apicolaterally acute (figs 29, 66)........................................................................7 [In some species with strongly vorticose elytral sculpture ( S. scopulus View in CoL -group) this character may be dubious: see below]

5 Paraglossae oval (fig. 15). Paratergites mostly less broad than antennal segment 2. Species of various lengths...................................................................................................6

- Paraglossae coniform (fig. 14). Paratergites distinctly broader than antennal segment, 2. Large species, FB>2.6mm............................................................... S. stigmaticus View in CoL -group

6: Median lobe without expulsion hooks or expulsion clasp (e. g. figs 307, 310, 311, 315); some setae of parameres often with a rough surface (figs 313, 314).: Spermatheca weakly sclerotized, very small, indistinct (fig. 308). Abdominal margination narrow, paratergites less broad than antennal segment 2, nearly impunctate, or absent (margination reduced to a simple line)............................... S. tenuimarginalis View in CoL -group

-: Median lobe with expulsion hooks and/or expulsion clasp (can be very small, e. g. in S. dapitananus BERNHAUER View in CoL , S. gardneri CAMERON View in CoL (fig. 301) and in S. montivagans CAMERON View in CoL ); surface of parameral setae smooth. Abdominal margination mostly as broad as antennal segment 2, mostly punctate ............................................ S. virgula View in CoL -group

7 Paraglossae coniform (fig. 14) ............................................................ S. abdominalis View in CoL -group (note: species of the S. stigmaticus- group also have coniform paraglossae)

- Paraglossae otherwise (mostly oval)...................................................................................8

8 Species with spotted elytra (1 exception: S. solstitialis ZHENG View in CoL ). Species of different phylogenetic groups ............................................................................................................9

- Species with non-spotted elytra.........................................................................................10

9: S proximately with a rattlesnake’s tail-like duct (fig. 403)............. S. tenuimargo View in CoL -group

-: S proximately with a swollen chamber (fig. 309)............................... S. indubius View in CoL -group

10 Paratergites in horizontal position.....................................................................................11

- Paratergites declining ventrad or only line-like.................................................................14

11 Greenish metallic, paratergites in horizontal position, broader than antennal segment 2, tarsi shorter (length of metatarsi: length of metatibiae  0,85) ............... S. viridanus View in CoL -group

- Blackish or with a bronze tint...........................................................................................12

12 Posterior margin of frons straight or slightly concave (figs 4, 5)......................................13

- Posterior margin of frons strongly concave (fig. 407 ....................... S. perplexus- group p.p.

13 Blackish, elytra strongly vorticose.: Spermathecal duct shorter, (mostly) bag-like swollen before collum (fig. 408) .............................................................. S. scopulus View in CoL -group

- Metallic with bronze tint.: Spermathecal duct longer, thin and multiply coiled, not swollen before collum (figs 278, 406).................................................... S. impressus View in CoL -group

14 Larger species, FB 2.8mm. Elytral sculpture strongly vorticose...... S. rugosipennis View in CoL -group

- Smaller species, FB 2.5mm. Elytral sculpture not strongly vorticose.............................15

15 Apicolateral tooth of sternum 9 not claw-shaped and not directed ventrad. Paraglossae oval......16

- Apicolateral tooth of sternum 9 claw-shaped, directed ventrad (fig. 400). Paraglossae silaniform (figs 16, 17, 376).................................................................... S. pilicornis View in CoL -group

16 One species from Thailand.: Aedoeagus ( Abb. 412 View Abb ). 3.0-4.0mm (FB 2.1mm).................. ............................................................................................................. S. falcipenis PUTHZ View in CoL

- Species from other countries.............................................................................................17

17 Species from Taiwan...................................................................... S. cephalotes View in CoL -group p.p.

- Species from S-India or Sri Lanka..................................................... S. brachypterus View in CoL group

C. Key to the Oriental species of the genus Stenus with unmargined abdomen and bilobed tarsi

(note: unmargined abdomen = no paratergites and no separation line on segments 4-6)

1 Tergite 10 with special characters (bicuspid, bifurcate, anchoriform: figs 167-191) [exceptions: S. aericeps BERNHAUER View in CoL (fig. 187), S. pernix L. BENICK View in CoL (fig. 168), S. scitulus L. BENICK View in CoL (fig. 169), with a very broad frons: HW: gDE <1.80].: Legs without any special sexual characters. FB 1.3- 2,2mm ............................... S. rorellus View in CoL -group (revision of the group see below)

- Tergite 10 otherwise............................................................................................................2

2 Tergite 10 apicolaterally roundly projecting, concave apicomedially (figs 48, 49). Frons comparatively narrow, pronotum slightly longer than broad. FB 1.9-2.1mm ............. ............................................................................................................... S. wasmanni View in CoL -group

- Tergite 10 rounded apically................................................................................................3

3 Tergite 8 with a posteromedian patch of dense shiny setae (figs 9, 12, 13)........................4

- Tergite 8 without a posteromedian patch of dense shiny setae...........................................8

4 Sternum 9 with a very long apicolateral tooth (figs 28, 57) ...................... S. bispinus View in CoL -group

- Sternum 9 otherwise............................................................................................................5

5 Sternum 9 of (fig. 61) [of serrate]....................... ( S. pulcher View in CoL -group). S. changi PUTHZ View in CoL

- Sternum 9 of otherwise...................................................................................................6

6 Sternum 9 with a moderately long apicolateral tooth (fig. 53).......... S. cicindeloides View in CoL -group

- Sternum 9 serrate apicolaterally or with a very short apicolateral projection (fig. 56).......7

7 Antennae very slender, when reflexed extending beyond the posterior margin of pronotum, penultimate segments much longer than broad. Antennae and legs yellowish .... ............................................................................................................. S. coelogaster View in CoL -group

- Antennae short, when reflexed extending to about middle of pronotum, penultimate segments broader than long. Antennae and legs predominantly dark ................................... .......................................................................................................... S. periscelidifer- group

8 Sternum 9 serrate apically (figs 63, 65, 225).......................................................................9

- Sternum 9 acute apicolaterally (figs 33, 34, 227)..............................................................11

9 Sternum 9 with an apicolateral brush about half as long as the sternum (fig. 225). Pronotum comparatively short (PL: PW <1.3). Smaller species: FB 1.8-2.3mm.................. .................................................................................................................. S. leptopus View in CoL -group

- Sternum 9 not or only with a short apicolateral brush. Pronotum short or longer. On average larger species.......................................................................................................10

10 Pronotum much longer than broad (PL: PW>1.3).: Legs simple. E: Median lobe in most cases without a flagellum ......................................................... S. cylindricollis View in CoL -group

- Pronotum shorter (PL: PW <1.25).: Legs often with special sexual characters. E: Median lobe often with a flagellum ..................................................... S. pulcher View in CoL -group pp.

11 Abdomen with conspicuously long and erect setae (figs 6, 7) ................ S. cirrus- group pp.

- Abdomen not with conspicuously long and erect setae.....................................................12

12 Sternum 9 with a very long apicolateral tooth (fig. 30).....................................................13

- Sternum 9 with a shorter apicolateral tooth (figs 33, 34)..................................................14

13 Larger species: FB>2.5mm. Frons concave, without distinct shiny plaques......................... ..... S. azurescens View in CoL -group (probably a mixed group of different phylogenetic relationships)

- Smaller species: FB 2.2mm. Frons convex, with distinct shiny plaques. S. piliferus View in CoL -group

14 Larger species, FB 1.9mm..............................................................................................15

- Smaller species: FB <1.9mm ................................................................ S. cephalotes- group (mixed group, probably phylogenetically belonging to species groups with margined abdomen)

15 Sternum 9 apicolaterally acutely ± prominent (figs 52, 54)..............................................16

- Sternum 9 apicolaterally less and roundly prominent (fig. 51) ................. S. lampros- group

16 Smaller species, FB <2.2mm.: No sclerotized spermatheca ............ S. pulcher- group pp.

- Larger species, FB>2.6mm.: With a sclerotized spermatheca (figs 263, 274)................. ................................................................................................................... S. guttalis- group

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

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