Patrera borjai, Martínez & Brescovit & Villarreal & Oliveira, 2021

Martínez, Leonel, Brescovit, Antonio D., Villarreal, Eduardo & Oliveira, Luiz Fernando M., 2021, An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia, Zootaxa 4914 (1), pp. 1-64 : 13-16

Patrera borjai new species

Figures 6A–G ; 31

Type material. Male holotype from Rivera Río Ñambí (1°14′44.88″N; 78°6′54.36″W), 1440m, Altaquer, Nariño, Colombia, 10.X.2009, E. Flórez & Estudiantes de Taxonomía Animal leg. (ICN-Ar-6074). Paratypes: female from Reserva Natural Rio Ñambi (1.28°N; 78.7°W), Vereda Altaquer 1335m., Barbacoas, Nariño, Colombia, 02- 06.VII.2017, D. Martinez et al. leg. ( IBSP 237017 ). One male from surroundings of the entrance of the Reserva Natural Río Ñambí (1°17′4.38″N; 78°5′18.24″W), 1179m, Vereda Altaquer , Barbacoas , Nariño, Colombia, 5.VII.2017, W. Galvis leg. (ICN-Ar-9673); one male from surroundings Cuadrante , Reserva Natural Río Ñambí (1°17′4.38″N; 78°5′18.24″W), 1262m, Vereda Altaquer , Barbacoas , Nariño, Colombia, 6.VII.2017, W. Galvis leg (ICN-Ar-9660) .

Other material examined. COLOMBIA, Nariño: Altaquer, Rivera Río Ñambí (1°14′45.22″N; 78°6′54.71″W), 1450m, ♁2, 14.IX.2014, D. Martínez & D. Luna leg. (ICN-Ar-10605) . Same locality (1.28°N; 78.7°W), 1335m, ♁1, ♀ 1, 02-06.VII.2017, D. Martinez et al. leg. ( IBSP 237018 ) .

Etymology. The specific epithet is a patronym given in honor of Rafael Borja, who has greatly contributed to ecological studies of Neotropical birds, for his support and advice to the first author.

Diagnosis. Males of Patrera borjai n. sp. resemble those of P. quillacinga n. sp. and P. perafani n. sp. by their prominent distal projection of the subtegulum, and a long embolus with narrow base no denticles ( Figs 11 C–E, 9C–E), but can be diagnosed by the embolus with wider base and reduced basal projections (very narrow and no projections in P. quillacinga and P. perafani ), and short, ventrally projected, apically truncated retrolateral tibial apophysis (shorter and apically quadrangular in P. quillacinga , and longer and apically sharp in P. perafani ) ( Fig. 6C ̅E). Females resemble those of P. quillacinga by having thin lateral borders, but can be distinguished by lateral borders separated in the medial region, forming two long and narrow atrial cavites, rounded and smaller spermathecae (larger in P. quillacinga ), and hood with rounded posterior edge ( Fig. 6F ̅G).

Description. Male (Holotype, ICN-Ar-6074). Carapace brownish, darker in the cephalic region ( Fig. 6A ). Chelicerae brown. Labium and endites yellow. Sternum yellow. Legs yellow, metatarsi and tarsi darkest. Abdomen dorsally orange; ventrally yellow. Spinnerets yellow ( Fig. 6A ). Total length 7.93, carapace length 3.88, width 2.94, high 1.21. Clypeus height 0.13. Eye diameters and interdistances:AME 0.11, ALE 0.2, PME 0.21, PLE 0.19; AME– AME 0.29, AME–ALE 0.4, PME–PME 0.53, PME–PLE 0.54, ALE–PLE 0.44. Chelicerae 1.71 long, four promarginal teeth; five retromarginal denticles. Leg measurements: leg I—femur 3.78/ patella 0.98/ tibia 4.14/ metatarsus 1.51/ tarsus 1.22/ total 11.63; II—3.67/ 1.09/ 3.98/ 2.63/ 1.45/ 12.82; III—2.8/ 0.77/ 2.3/ 2.59/ 1.24/ 9.70; IV—3.17/ 1.08/ 3.59/ 2.58/ 1.48/ 11.59. Leg spination: I—metatarsus v2-0-2; II—metatarsus v2-1-1; III—tibia d1-0-1, p1-1-1, r1-1-1, metatarsus p1-2-2, r1-1-2; IV—tibia d1-0-1, metatarsus d0-1-0, p1-1-2, r1-1-2. Abdomen: length 4.4, epigastric furrow 0.96 from tracheal spiracle, spiracle 1.56 from base of spinnerets. Palp: retrolateral tibial apophysis short and ventrally projected; cymbium longer than tibia; subtegulum longer than wide with prominent and apically subrounded distal projection; tegulum longer than wide; median apophysis hook-shaped, long and medially situated; embolus long with wide base, apically filiform and basally inserted on the tegulum ( Fig. 6C ̅E).

Female (Paratype, IBSP 237017). Coloration as in the male ( Fig. 6B ). Legs yellow. Total length 9.8, carapace length 3.8, width 2.3. Clypeus height 0.14. Eye diameters and interdistances: AME 0.10, ALE 0.18, PME 0.18, PLE 0.18; AME–AME 0.02, AME–ALE 0.08, PME–PME 0.14, PME–PLE 0.18, ALE–PLE 0.08. Chelicerae 1.5 long, with four promarginal teeth and five retromarginal denticles. Leg measurements: leg I—femur 3.2/ patella 1.1/ tibia 3.3/ metatarsus 2.5/ tarsus 1.55/ total 11.65; II—3.2/ 1.1/ 3.0/ 2.4/ 1.3/ 11.0; III—2.5/ 1.0/ 1.4/ 2.4/ 0.9/ 8.2; IV—3.25/ 1.3/ 2.6/ 3.5/ 1.2/ 11.85. Leg spination: I—metatarsus v2-2-0, p0, r0; II—metatarsus=I; III—tibia v2-2-2, p1-1d-0-1, r0-1-1, metatarsus v2-2-2, p1-1-1, r1-1-1; IV—tibia v2-2-2, p0-1-1, r1-1-1, metatarsus v2-2-2, p1-1-1, r1-1-1. Abdomen: length 5.9, epigastric furrow 1.8 from tracheal spiracle, spiracle 2.4 from base of spinnerets. Epigynum: hood triangular, subrounded in the posterior edge; lateral borders sclerotized, parallel, long, wide and apically rounded; atrial cavities long and narrow; internally with copulatory ducts long and thin; seminal receptacles very small and posteriorly situated on the spermathecae; spermathecae large, rounded and posteromedially positioned; fertilization ducts almost as long as spermathecae ( Fig. 6 F–G).

Variation. Males (n=5): total length: 7.55–8.66; carapace length: 3.73–4.45; femur I length: 3.72–3.86.

Distribution. Only known from Nariño department ( Fig. 31 ).

Figures

FIGURE 6A–G. Patrera borjai n. sp. Male (ICN-Ar-6074): A habitus, dorsal view; B left palp, retroventral view; C palp, ventral view; D palp, retrolateral view. Female (IBSP 237017): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Scale bars: A, B: 2mm; C: 1mm; D–E: 0.5mm; F–G: 0.2mm.

FIGURE 11A–G. Patrera quillacinga n. sp. Male (ICN-Ar-9676):A habitus, dorsal view; C left palp, retroventral view; D palp, ventral view; E palp, retrolateral view. Female (IBSP 213421): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Scale bars: A–B: 2mm; C: 0.7mm; D–E: 0.5mm; F: 0.5mm; G: 0.2mm.

FIGURE 31. Distribution of species of the fulvastra group in Colombia.