Gymnetina Casey, 1915

Ratcliffe, Brett C. & Warner, William B., 2011, A revision of the genus Gymnetina Casey, 1915 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini), Insecta Mundi 2011 (173), pp. 1-28 : 4

Key to the Species of Adult Gymnetina Casey, 1915

1. Pronotum and elytra with large, dense, mostly confluent punctures. Dorsal surface completely black. Guatemala....................... G. grossepunctata Ratcliffe and Warner , new species

– Pronotum with minute to small punctures; if with punctures moderate in size then also with cretaceous bands along lateral margins. Elytra sparsely to moderately punctate on disc with punctures becoming denser on sides but never completely, coarsely punctate; cretaceous marks present or not. Mexico and USA............................................................................................... 2

2(1). Elytron with 2 large, round, cretaceous spots in transverse row just behind middle ( Fig. 11a ); rarely, spots reduced to a single, submarginal spot ( Fig. 11b ) or spots fused ( Fig. 11c ). Male with protibial teeth reduced to obsolete. Parameres (ventral view) lacking prominent tooth at base ( Fig. 14 ) ............................................................................................................................ 3

– Elytron with slender or broad, cretaceous band just behind middle ( Fig. 25a ); rarely broken into 2 irregular spots ( Fig. 25b ) or with small, cretaceous flecks or without cretaceous marks. Male protibial teeth not reduced. Parameres (ventral view) with distinct tooth at base ( Fig. 28 ) .. 4

3(2). Pronotum with broad, cretaceous band along lateral margins ( Fig. 11 ). Mesepimeron usually with cretaceous spot. Pygidium with large, cretaceous spot either side of middle. SE Arizona, extreme SW New Mexico, and Chihuahua and Durango, Mexico.............................................. ............................................................................................ G. cretacea cretacea (LeConte)

– Pronotum with slender, partial, cretaceous band along lateral margins ( Fig. 15 ) or lacking cretaceous marks altogether. Mesepimeron never with cretaceous spot. Pygidium with or without cretaceous spot either side of middle. Doña Ana Co. New Mexico .............................................. ...................................... G. cretacea sundbergi Warner and Ratcliffe , new subspecies

4(2). Pronotum with broad, cretaceous band along lateral margins ( Fig. 25 ). Elytron with broad, cretaceous, transverse band just behind middle. SE Arizona, SW New Mexico, and Chihuahua and Sonora, Mexico................................. G. howdeni Warner and Ratcliffe , new species

– Pronotum with slender, cretaceous band along lateral margins ( Fig. 1 ) or without any cretaceous marks. Elytron with slender, cretaceous, transverse band just behind middle or with a small, cretaceous fleck near lateral margin (rarely with 2 small flecks traversing elytron) ............. 5

5(4). Pygidium without cretaceous spots ( Fig. 7 ). Central Arizona and New Mexico ............................ .................................................................. G. borealis Warner and Ratcliffe new species

– Pygidium with cretaceous spot either side of middle. Durango to Oaxaca, Mexico.................... 6

6(5). Mesepimeron usually with cretaceous spot. Elytron with slender, sinuate, transverse, cretaceous band just behind middle (band rarely broken). Males with cretaceous spot on posterolateral margin of sternites 1-4 (variably reduced) AND on anterior margin of sternites 2-5 either side of middle (variably reduced, but always with at least 1 spot). Southern Mexico ........................... ......................................................................................................... G. alboscripta (Janson)

– Mesepimeron without cretaceous spot. Elytron with single, small, cretaceous spot near lateral margin just behind middle. Males with cretaceous spot on posterolateral margin of sternites 1-4 (variably reduced) but never with second set of cretaceous marks on sternites 2-5 either side of middle. Mexico.............................................................................................. G. salicis (Bates)

Figures

Figures 11-14. Gymnetina cretacea cretacea: 11a-c) Dorsal views, showing variations of cretaceous pattern on pronotum, mesepimeron, and elytra as well as distinctive punctation on pronotum. 12) Lateral view showing cretaceous metepisternum, pygidial spot, flat mesometasternal process, single row of cretaceous spots on sides of abdomen and distinct concavity of male abdomen. 13-14) Dorsal and ventral views, respectively, of parameres. Note absence of tooth at base of paramere on ventral side (arrow).

Figures 25-28. Gymnetina howdeni: 25a-b) Dorsal views, showing variation of cretaceous pattern on pronotum and elytra as well as sparse, small punctures on pronotal disc. 26) Oblique ventral view showing double row of cretaceous spots on sides of male abdomen, pygidial spot, and more broadly rounded apex of mesometasternal process. 27-28) Dorsal and ventral views, respectively, of parameres. Note distinct tooth at base of paramere on ventral side (arrow).

Figures 15-19. Gymnetina cretacea sundbergi: 15) Dorsal view, showing reduced cretaceous pattern on pronotum as well as distinctive punctation on pronotum. 16) Ventral view showing narrower apex of mesometasternal process and reduced cretaceous spots on sides of abdominal sternites. 17) Caudal view showing sculpturing and markings on pygidium. 18-19) Dorsal and ventral views, respectively, of parameres. Note absence of tooth at base of paramere on ventral side (arrow).

Figures 1-4. Gymnetina alboscripta: 1-2) Dorsal and ventral views, respectively. 3-4) Dorsal and ventral views, respectively, of parameres. Note distinct tooth at base of paramere on ventral side (arrow).

Figures 5-10. Gymnetina borealis: 5-6) Dorsal views, showing reduced cretaceous pattern on pronotum, mesepimeron, and elytra as well as sparse, small punctures on pronotal disc. 7) Caudal view showing sculpturing and absence of markings. 8) Oblique ventral view showing broader apex of mesometasternal process and reduced cretaceous spots on sides of abdomen. 9-10) Dorsal and ventral views, respectively, of parameres. Note distinct tooth at base of paramere on ventral side (arrow).