Lucanus kraatzi kraatzi Nagel, 1926

Zhan, Zhihong & Young, Daniel K., 2023, A taxonomic assessment and redefinition of the Lucanus fortunei species group in China (Coleoptera: Lucanidae: Lucaninae), Zoological Systematics 48 (4), pp. 279-360 : 336-337

publication ID	https://doi.org/ 10.11865/zs.2023401
publication LSID	lsid:zoobank.org:pub:46AA3E9C-C065-415D-946B-0813AF169B14
persistent identifier	https://treatment.plazi.org/id/A51987E0-7F6D-FFA6-FF02-AC25FD09FECD
treatment provided by	Felipe
scientific name	Lucanus kraatzi kraatzi Nagel, 1926
status

Treatment

Lucanus kraatzi kraatzi Nagel, 1926 View in CoL ( Figs 201–208 View Figures 201–208 , 271 View Figures 260–275 , 303 View Figures 276–307 , 331 View Figures 324–335 , 359 View Figures 336–363 , 442–444 View Figures 436–456 )

Lucanus kraatzi Nagel, 1926: 116 View in CoL ; Didier, 1927: 202; Didier & Séguy, 1952: plate 17, fig. 4; Mizunuma & Nagai, 1994: 216, plate 16, figs 109-1–3; Zilioli, 2003: fig. 2.

Lucanus kraatzi kraatzi Ikeda, 1997: 14–16 View in CoL , figs 5–6. Material examined. [5♂]: CHINA: Sichuan Province, Liangshan Region // Da-Liang-Shan // [dz山彝Dzss治州, kdz

山]; Alt. 1750m; 30 August 2019 // Zihao Yang leg.; [2nd label]: 27.8848°N; 102.2622°E Adult // collected on host plant// Cyclobalanopsis glauca View in CoL // (Thunberg) Oersted; [3rd label]: Lucanus View in CoL // kraatzi kraatzi View in CoL // Nagel ♂ // det. Z. Zhan; [5♂]: CHINA: Yunnan Province, Honghe Pref., // Pingbian County, Da-Wei-Shan Mt., // [Ḧ河屏®, k围山]; Alt. 2000m; 12 Septemer. 2021 // Xv Yan leg.; [2nd label]: 22.9896°N; 103.6736°E Adult // collected by Light-trap//; [3rd label]: Lucanus View in CoL // kraatzi kraatzi View in CoL // Nagel ♂ // det. Z. Zhan; [1♀]: CHINA: Guizhou Province, Liupanshui City // Niang-Niang-Shan Mt., // [六#水市, OiOi山]; Alt. 1650m; 20 August. 2017 // Zhihong Zhan leg.; [2nd label]: 26.5925°N; 104.8303°E Adult // Light-trap collected; [3rd label]: Lucanus View in CoL // kraatzi kraatzi View in CoL // Nagel ♀ // det. Z. Zhan.

Body length 46.2–68.4 mm (♂); 33.5 mm (♀).

Description. Major male. Body brownish to black, surface metallic, covered with vestiture of short, yellowish setae. Head, including widest region of mandibular bases, 1.65× width of pronotum and elytra. Head brownish to black, surface covered with vestiture of short yellowish setae; mandibles strongly incurved at distal 1/3, straight along midlength and apex; major inner mandibular tooth located approximately at middle of each mandible, sharply triangular and well developed;>8 small inner mandibular teeth located above and below major mandibular tooth. Apical mandibular forks developed, upper branches of apical fork 1.45× length of lower apical fork branches; anterolateral ridges of head strongly convex, posterolateral margins of head widened, significantly wider than anterolateral ridges; clypeolabrum stout, trapezoidal; labrum triangular, protruding along midlength. Thorax brownish to black, concolorous with head, surface setose; convex at apex and base, significantly protruding at middle; 0.6× width of head and mandibles; prothoracic tibiae reddish to brownish, each bearing four slender spines from apex to midlength; mesothoracic tibiae reddish to brownish, each bearing 2–3 spines along midlength; metathoracic tibiae reddish, each bearing a single poorly developed spine distally. Elytra brownish to black, usually concolorous with head and pronotum, less setose and metallic along suture; elytral margins slender, widest at apical 1/3, strongly narrowed at basal 1/3. Median male as major male except: head with posterolateral margins weakly widened. Minor male as major and median males except: head with posterolateral margins subparallel with anterior margins. Genitalia. Basal piece wide and short, 0.5× length of entire aedeagus; apical flagellum weakly enlarged, flagellum 0.45× length of entire aedeagus; aedeagus stout, incurved at basal 1/3, triangular distally.

Female. Body black, surface more setose than that of male, metallic; body stout, always smaller than major and median males, sometimes larger than or subequal in length to that of minor male. Head black, surface punctate, with metallic luster; mandibles straight along midlength and apical 1/3; each mandibular tooth well developed, asymmetrical; right mandibular tooth triangular, left mandibular tooth weakly convex along midlength; preocular and postocular margins strongly developed, forming acute angles, and protruding anteriorly; clypeolabrum single-pointed. Thorax pronotum always concolorous with head, surface shinning, covered with vestiture of yellowish setae; anterior aspect of pronotum weakly enlarged, subequal in width to that of posterolateral pronotum; prothoracic tibiae slender and black, each bearing four spines from anterior to midlength; distal two spines sharp, well developed; prothoracic tibiae 1.3× width of mesothoracic and metathoracic tibiae; mesothoracic tibiae black, each bearing two spines along midlength; metathoracic tibiae black, each bearing a single spine at basal 1/3. Elytra black, with metallic luster, concolorous with head and pronotum; oval, widest at apical 1/3, strongly narrow at basal 1/3.

Distribution. China (Sichuan, Guizhou, Yunnan).

Phenology. Adults of L. kraatzi kraatzi were collected from late August to mid-September.

Natural history. Typical plants for this species are Cyclobalanopsis glauca , with height usually less than 2 m. During this study, some males of L. kraatzi kraatzi were also observed active on grassland. Adult longevity is around 25–30 days.

Collecting methods. Hand and light-trap collected.