Thorodiplosis Felt

Kolesik, Peter & Gagné, Raymond J., 2020, A review of the gall midges (Diptera: Cecidomyiidae) of Indonesia: taxonomy, biology and adult key to genera, Zootaxa 4847 (1), pp. 1-82 : 69

Thorodiplosis Felt

[ Figs 40 a–j]

Thorodiplosis Felt, 1921d: 89 .

Type species: Thorodiplosis impatientis Felt , by original designation.

Description. This monotypic genus was erected for two males and two females based on rather broad characteristics: very long and narrow wings, elongate antennae, simple tarsal claws curved at almost right angle, short ovipositor with bilobed cercus, and 4-segmented palpus ( Felt 1921d). The types are uncleared and partially shriveled but all characters are recognizable, except for the male circumfila that were missing already when Felt (1921b) examined them. Wing [ Fig. 40g ] with R 5 bent distally, joining C posteriad of wing apex; C broken at juncture with R 5; M 4 and Cu 1 forming fork; Rs not present. Head with occipital protuberance present. Palpi shriveled on all specimens but evidently four-segmented. Antennae: 12 flagellomeres, progressively shorter, necks bare of microtrichia; binodal in male, presumably trifilar, with long necks [ Fig. 40i ]; cylindrical in female, all except last with long necks, proximal flagellomeres with nodes 3x as long as wide, constricted at basal third, distal flagellomeres with nodes 2x as long as wide, progressively losing basal constriction, circumfila consisting of one uneven horizontal and two more distal vertical bands [ Figs 40c, d ]. Tarsal claws simple, narrow, bent at right angle at midlength, as long as empodia [ Fig. 40e ]. Male terminalia [ Figs 40h, j ]: gonocoxite narrow, with low, wide mesobasal lobe; gonostylus very long, narrow, widest at distal fourth, straight and setulose basally, curved and carinate beyond, with small, solid, tooth; cerci broad and concave apically, the mesal lobe of the concavity with single seta; hypoproct longer than cerci, narrow, incised apically, resulting lobes each with single seta apically; aedeagus longer than hypoproct, tapered to narrowly rounded apex, with asetose papillae distally. Female terminalia [ Fig. 40f ]: ovipositor short, cerci large, ovoid, setose mainly distally, each with pair of long, blunt apical sensory hairs; hypoproct with pair of setae.

Remarks. We place Thorodiplosis in the tribe Clinodiplosini because of the conformation of the male terminalia, particularly the apically concave cerci.

Figures

FIGURES 40a–j. Thorodiplosis impatientis: a–b, galls on Impatiens platypetala: a, galls indicated with arrows (Fig. 599 of DvL & DvLR 1918); b, Cianjur near Mt Gede (photo by Purnama Hidayat and Mahindra Dewi Nur Aisyah, 2018); c–g, female: c, 1st and 2nd flagellomeres; d, antenna; e, tarsal claw and empodium; f, cerci and hypoproct (lateral); g, posterior part of wing; hj, male: h, terminalia (dorsal); i, 6th flagellomere (circumfila not visible on types); j, aedeagus of both type specimens (dorsal).