Daviesia hakeoides Meisner (1844: 47)

Crisp, Michael D., Cayzer, Lindy, Chandler, Gregory T. & Cook, Lyn G., 2017, A monograph of Daviesia (Mirbelieae, Faboideae, Fabaceae), Phytotaxa 300 (1), pp. 448-450 : 448-450

91. Daviesia hakeoides Meisner (1844: 47) , Crisp (1987a: 250), Crisp (1995: 1196). Type: Syntypes are cited under the subspecies below. Lectotype (Crisp 1995: 1196): Preiss 1156 (LD); isolectotypes: GOET, K, MEL (2 sheets), MO, NY, P (2 sheets), W (2 sheets)

Intricate, many-stemmed shrubs, 0.3–1 m tall and 0.5–2 m broad, glabrous, glaucescent. Root anatomy in both subspecies with anomalous secondary thickening (cord roots). Branchlets spreading to ascending, terete, smooth when fresh, striate when dry. Phyllodes scattered, divaricate to erect, terete, pungent, inarticulate and continuous with the branchlet, 0–80 mm long and up to 1.5 mm broad at base, striate when dry, or often reduced to scales over most of plant. Juvenile phyllodes flat, narrow-oblong, 17–25 × ca. 5 mm (Crisp 6625). Unit inflorescences 1 per axil, condensed racemes with the lower flowers clustered due to condensation of the rachis, 2–6-flowered, enclosed at base by large, overlapping bracts; peduncle 1–1.5 mm long; rachis very short (<2 mm long); barren basal bracts

204 • Phytotaxa 300 (1) © 2017 Magnolia Press

CRISP ET AL.

A MONOGRAPH OF DAVIESIA

Phytotaxa 300 (1) © 2017 Magnolia Press • 205 forming an involucre, shell-shaped to oblong, ca. 1–2 mm long; subtending bracts ascending, spathulate, shell-shaped, striate, claw fused to pedicel for up to 1 mm, up to 3.5 mm long. Pedicels 1–6 mm long. Calyx 2–2.5 mm long including the ca. 0.5 mm receptacle; lobes very short (<0.25 mm long); upper 2 lobes ± united; lower 3 lobes acuminate, flared outwards then incurved at the tips; base of receptacle abruptly contracted to articulation with pedicel. Corolla : standard transversely broadly elliptic, emarginate, cordate, 4.5–6 × 5–7 mm including the ca. 1 mm claw, with a deep channel and 2 small calli towards the base of the lamina, yellow or orange with a dark red centre; wings obliquely obovate with a rounded and incurved apex that encloses the keel, auriculate, 5–5.5 × 2–2.5 mm including the 1–1.5 mm claw, dark red; keel half transversely broadly elliptic with an acute apex, slightly auriculate, saccate, ca. 5 × 1.75 mm including the 1.5–2 mm claw, dark red. Stamens strongly dimorphic: inner whorl of 5 with longer, slender, terete filaments and shorter, round anthers with confluent thecae; outer whorl of 5 with shorter, broader, compressed filaments and longer, oblong, 2-celled anthers; filaments cohering. Pod obliquely shallowly obtriangular, more or less beaked, somewhat turgid, 14–17 × 10–11 mm, thick-walled; upper suture ± sigmoid; lower suture forming an acute angle centrally and indented towards the apex. Seed with a continuous aril. ( Fig. 91 ).

Flowering period:— May to July. Fruiting period: August to November.

Distribution:— South-west Western Australia from north of Kalbarri southward to near Albany and inland in the northern, central and southern wheatbelt.

Affinity:— This species belongs to a group of closely related species with moderately enlarged, shell-shaped, usually striate bracts that are imbricate and cover the rachis ( Crisp 1982a; 1984), and subsp. hakeoides is most similar to D. debilior subsp. debilior , D. major and D. smithiorum (see discussion under subsp. hakeoides ). Daviesia hakeoides subsp. subnuda has been confused with D. debilior , D. gracilis and D. triflora (see discussion under subsp. subnuda ).

Figures

FIGURE 91. Daviesia hakeoides subsp. hakeoides. A. Flowering branchlet (typical form with ascending phyllodes). B. Fruiting branchlet (northern form with divaricate phyllodes). C. Basal linear phyllode. D. Inflorescence. Daviesia hakeoides subsp. subnuda. E. Flowering branchlet. F. Pod. A, D from Crisp 6620; B from Crisp 6263; C from Crisp 6622; E from Crisp 6511; F from Crisp 6321. Drawn by A.L. Prowse.