Daviesia nova-anglica Crisp (1990: 245)

Crisp, Michael D., Cayzer, Lindy, Chandler, Gregory T. & Cook, Lyn G., 2017, A monograph of Daviesia (Mirbelieae, Faboideae, Fabaceae), Phytotaxa 300 (1), pp. 448-450 : 448-450

61. Daviesia nova-anglica Crisp (1990: 245) , Crisp (2002: 526). Type: New South Wales, Northern Tablelands, ca. 23 km E of Tenterfield, 8.5 km from Timbarra along road to Poverty Point, 29°04’S, 152°15’E, M.D. Crisp 7336 & I.R. Telford, 29 September 1984. Holotype: CBG; isotypes: BRI, K, MEL, MO, NSW

Shrubs 0.5–1(–2) m high, hispid on branchlets, minutely so on midrib and margins of phyllodes. Root anatomy unknown. Branchlets arching, terete, ribbed, hispid. Phyllodes fairly crowded, ± divaricate, ovate or narrowly so,

142 • Phytotaxa 300 (1) © 2017 Magnolia Press

CRISP ET AL.

rarely broad, acute or acuminate at apex, pungent, ± recurved at margins, subcordate to rounded at base, articulate, 5–14 × 2.5–7 mm, green; midrib prominent above; venation obscure. Unit inflorescences 1 per axil, flowers solitary or 2 in umbels; peduncle 0.8–4.0 mm long; barren basal bracts ascending, oblong; subtending bracts ascending, oblong, 1–1.5 mm long, 0.5–0.75 mm broad. Pedicels 3–6.25 mm long. Calyx 2.5–3 mm long including 0.5–0.8 mm receptacle; lobes ca. 0.75 mm long; upper 2 united in a truncate emarginate lip; lower 3 broadly triangular. Corolla : standard depressed-ovate, emarginate, scarcely auriculate and with slight calli at the base, 6–7 × 7.5–8.5 mm including the 1–1.5 mm claw, yellow with maroon infusion surrounding an oblong or bilobed central yellow spot; wings obovate, rounded and incurved at apex, auriculate at the base, 5.5–6.5 × 2.5–3 mm including the ca. 1.5 mm claw, yellow at the tips and margins with maroon infusion towards the base; keel half very broadly ovate, acute, auriculate, saccate, ca. 5 × 2–2.5 mm including the ca. 1.5 mm claw, maroon. Stamens dimorphic though filaments are uniform in length; inner whorl of 5 with terete filaments and versatile, round, smaller anthers with confluent thecae; outer whorl of 5 with compressed filaments and larger, basifixed, 2-celled anthers; filaments

A MONOGRAPH OF DAVIESIA

Phytotaxa 300 (1) © 2017 Magnolia Press • 143 free. Pod obliquely shallowly obtriangular, acute, compressed, 6.5–8 × 5–6 mm; upper suture sigmoid; lower suture acute. Seed not seen. ( Fig. 61 ).

Chromosome number:— n = 9, 2n = 18 (voucher Sands 628.2.12, cited under D. squarrosa var. squarrosa by Sands 1975).

Flowering period:— August to October. Fruiting period: November.

Distribution:— Known only from New South Wales, where it occurs mainly along the eastern escarpment of the Northern Tablelands from near Tenterfield in the north to the headwaters of the Hastings River in the south; also recorded near Warialda on the North-west Slopes. This species possibly occurs in Queensland because one population (Crisp 7310) is only 1 km from the state border.

Habitat:— Grows in sandy soils, usually derived from granite, at elevations of 500–1000 m. Associated vegetation is open forest dominated by eucalypts, with a sclerophyll shrub understorey; less commonly occurring in woodland or heath.

Selected specimens (26 examined):— NEW SOUTH WALES. North-west Slopes: 3 km from Warialda, ca. 29°03’S, 150°34’E, Anon., 15 October 1960 ( NE 32450 ). Northern Tablelands: 1 km along road to Stanthorpe ( Qld ) from turn-off 0.5 km N of Wilsons Downfall on Mt Lindesay Hwy , 28°41’S, 152°04’E, M . D. Crisp 7310 & I. R . Telford, 28 September 1984 ( BRI, CBG, NSW); Gibraltar Range , 1 km S of Boundary Trig, 29°33’S, 152°16’E, M . D. Crisp 7369 & I. R . Telford, 29 September 1984 ( CBG, NSW); Werrikimbe Trail, Werrikimbe National Park , 31°10’S, 152°16’E, P . Gilmour 5909, 14 October 1986 ( CBG); Gibraltar Range, J. W . Green 2734, 25 November 1960 ( NE); Gibraltar Range State Forest , 29°35’S, 152°13’E, V. E . Sands 628.2.12, 2 August 1962 ( SYD); Forbes River, Mt Boss State Forest , 51 km NW of Wauchope, 31°06’S, 152°21’E, H . Streimann 8212, 20 October 1978 ( CBG, K, NSW). CULTIVATED. Australian National Botanic Gardens, section 55, no. 8410698 (ex Gibraltar Range ), M . D. Crisp 8247, 20 October 1988 ( CBG, NSW) .

Affinity:— Specimens of Daviesia nova-anglica were once referred to D. squarrosa var. squarrosa (= D. squarrosa ). Daviesia squarrosa may be distinguished most readily by its smaller upper bracts (0.5–0.8 mm long and 0.3–0.4 mm broad). Phyllode shape also differs in these species, although the distinction is more subtle. In D. squarrosa , the phyllodes are basically heart-shaped with a pronounced cuspidate apex 3–5 mm long, whereas in D. nova-anglica , they are basically ovate with a shorter, acuminate apex up to 4 mm long. Daviesia villifera and D. pubigera differ from both these species in being hispid all over the phyllodes, and D. villifera is further differentiated by the hooked, beaked keel-petal.

Figures

FIGURE 61. Daviesia nova-anglica. A. Branchlet late in flowering. B. Phyllode, abaxial view. C. Phyllode, adaxial view. D. Inflorescence (1-flowered). E. Calyx opened out, upper lobes at left. F. Standard. G. Wing. H. Keel. I. Pod. A–C from Crisp 8247; D– H from Crisp 7336 (type); I from Green 2734. Drawn by J. Marsh. Adapted from Crisp (1990) with permission from CSIRO Publishing.