Ptychoderes mixtus Jekel, 1855

Mermudes, José Ricardo M. & Napp, Dilma Solange, 2006, Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini), Zootaxa 1182 (1), pp. 1-130 : 44-50

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https://doi.org/ 10.11646/zootaxa.1182.1.1

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scientific name

Ptychoderes mixtus Jekel, 1855
status

 

Ptychoderes mixtus Jekel, 1855

Figs. 71 View FIGURES 70–84 , 85, 99 View FIGURES 85–112 , 113 View FIGURES 113–126 , 130–133 View FIGURES 127–141 , 185–188 View FIGURES 177–195 , 199, 201 View FIGURES 196–201 , 219 View FIGURES 215–220 , 232, 236 View FIGURES 228–238 , 266 View FIGURES 265–266. 265

Ptychoderes mixtus Jekel, 1855: 57 ; Gemminger & Harold, 1872: 2722 (cat.); Bovie, 1906: 220

(cat.); Jordan, 1906: 303, t. 10, fig. 3; Wolfrum, 1929: 6 (cat.); Rheinheimer, 2004: 10 (cat.). Ptychoderes mixta [sic]; Blackwelder, 1947: 765 (cat.).

Male. Vestiture ( Fig. 219 View FIGURES 215–220 ): pale dorsal vitta with dense whitish to clay­whitish scales, intermingled with brownish scales, decumbent and distinctly denser on rostrum, head, elytral interstriae 1, 2 and 4, apical declivity and epipleura. Pygidium with predominance of brown scales. Ventral surface and legs with appressed, dense yellowish­white scales; metasternum and ventrites medially densely pilose; sides of prosternum, below lateral carina, with a dark vitta of blackish­brown scales narrowed posteriorly towards lateral carina and narrowly divided by whitish scales runnin along pleural suture. Sides of mesosternum with a narrow glabrous vitta. Mesepimera, episterna, sides of metasternum and ventrites with coarse setigerous punctures, each provided with a brownish scale. Outer surface of meso­ and metafemora with a strip of blackish­brown scales.

Rostrum 1.3–1.8 times as long as wide across base; apical notch deep, narrow. Distance between scrobe and eyes as long as maximum width of eye. Sides barely swollen, with two straight, thickened carinae; near the basal margin of scrobe either with coarse, confluent punctures or with an impressed area, carinae vanishing. Scrobes careniform at dorsal and basal margins; extension vanishing with a deep distal pit. Mentum with sparse, moderately coarse punctures; groove of apical margin vestigial. Gula with sparse, fine punctures. Frons with prominent carinae.

Antennae ( Fig. 219 View FIGURES 215–220 ) from short, reaching elytral basal third, to slightly longer than body, exceeding elytral apices by club length; segments II–VII increasing in length; VIII–IX decreasing; X one third shorter than XI (II = 0.4–1.2; III = 0.6–2.2; IV = 0.6–2.8; V = 0.6–3.2; VI = 0.5–3.5; VII = 0.5–3.7; VIII = 0.4–3.3; IX = 0.6–2.8; X = 0.3–0.8; XI = 0.5–1.2).

Prothorax ( Fig. 219 View FIGURES 215–220 ) as long as or longer than wide; sides converging anteriorly from antebasal carina. Pronotal depression deep with a rounded, weakly prominent, median tubercle not reaching margins of depression. Pronotum transversely rugose throughout, more so on disk and at sides. Antebasal carina narrowly interrupted at middle (subequal to width of carina), oblique towards lateral carina. Lateral carina strongly sinuous. Secondary carina widely interrupted at middle, almost reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Disk of prosternum raised, with irregularly dense, deep, coarse punctures; antecoxal furrow deep, straight, narrow and entire. Mesosternum with sparse, shallow, moderately coarse punctures, and a deep, irregularly transverse impression coarsely punctate at sides near intercoxal process. Intercoxal process of mesosternum feebly impressed at apex, not swollen at lateral and apical margins, without lateroapical projections. Elytra 2.1 times as long as wide across humeri, weakly depressed at basal half; interstria 1 raised for the entire length, continuing with apical margin; basal gibbosities weakly prominent; surface rugose from base to behind middle, more so at base and near suture. Sides subparallel, convergent at apical 1/4. Anteapical tubercles ( Fig. 232 View FIGURES 228–238 ) rounded, from developed to weakly prominent, formed by the swollen interstriae 3–9. Wings without radial cell.

Abdomen. Ventrite I ( Fig. 236 View FIGURES 228–238 ) clothed with dense, erect, yellow pilosity, setiferous sex patch slightly developed, suboval, closer to apical margin; intercoxal process deeply grooved. Ventrite V feebly emarginate at apex. Pygidium feebly subconvex, with a short groove to fit elytra, apical margin truncated.

Terminalia. Tergite VIII ( Fig. 71 View FIGURES 70–84 ) slightly wider than long, subparallel at sides, almost rounded at apical margin; sternite VIII ( Fig. 85 View FIGURES 85–112 ) convergent at sides, apical margin emarginate forming large lobes, apodeme triangular, pigmented. Apodeme of sternite IX ( Fig. 99 View FIGURES 85–112 ) 3.6 times as long as arms. Tegmen ( Fig. 113 View FIGURES 113–126 ) rounded at apex. Aedeagus ( Figs. 130–133 View FIGURES 127–141 ) with arc between apodemes; tectum ( Fig. 130, 131 View FIGURES 127–141 ) straight at proximal margin, acuminate apically from base; pedon ( Figs. 130, 132 View FIGURES 127–141 ) with body slightly curved and thickened, apex subacuminate; apodemes oblique at proximal third. Internal sac ( Fig. 130 View FIGURES 127–141 ) almost as long as aedeagus, almost entirely spiculated, more so at middle; median sclerites strongly sclerotised ( Fig. 133 View FIGURES 127–141 ).

Female. Rostrum slightly shorter, at least 1.2 times as long as wide across base, not swollen at sides; extension of scrobe shallow. Antennae shorter, reaching the middle of prothorax; segments II–VII subequal in length, VIII and X shortest; club thickened, segments IX and XI distinctly longer than preceding (II = 0.3–0.6; III = 0.3–0.5; IV = 0.3–0.6; V = 0.2–0.4; VI = 0.2–0.4; VII = 0.2–0.4; VIII = 0.2–0.3; IX = 0.4–0.6; X = 0.2–0.3; XI = 0.4–0.5). Elytra slightly shorter and convex. Ventrite V with apical margin truncate. Terminalia ( Figs. 185–188 View FIGURES 177–195 ). Tergite VIII ( Fig. 185 View FIGURES 177–195 ) slightly longer than wide, sides divergent from anterior margin to middle, then convergent towards truncate apical margin; sternite VIII ( Fig. 186 View FIGURES 177–195 ) slightly shorter than apodeme, truncate at apex; ovipositor ( Figs. 187, 188 View FIGURES 177–195 ) with four teeth: two distal, conchoidal, developed and subequal, one proximal, dorsolateral, moderately developed, and one small proximal ventrolateral; median rods stout and rounded at proximal margin; median ventral lobe bilobated; bursa copulatrix with ventral lobe.

Measurements, in mm, male/female, respectively. CT = 9.8–18.5/9.0–18.3; CR = 1.3–3.4/1.2–2.2; LAR = 1.2–2.1/1.2–2.0; LBR = 1.0–1.8/0.9–1.7; LC = 1.6–2.8/0.8–1.3; DEO = 0.6–3.0/0.5–0.8; MLO = 0.8–3.0/0.8–1.3; CP = 2.5–5.0/2.3–4.3; LP = 2.5–4.7/ 2.3–4.2; CE = 6.3–12.5/5.8–11.3; LE = 3.0–5.8/4.5–5.2.

Type material

Syntypes, 1 male and 2 females ( MNHN) examined : 1 male [Coll. Chevr.]; 1 female [ Brésil 37, Chevr.]; 1 female [Chevr.]. The specimens have no individual labels, there is, however, a unique label at the bottom of the box, written: “Mixtus Jek., Ins. Saunders., Brazil, Cayen.” Type locality: Jekel (1855) studied specimens from Cayenne ( French Guiana) and Rio Negro ( Brazil) but did not mention the number of specimens .

Remarks

Three synapomorphies support P. mixtus as the sister group of the clade P. bivittatus +: tergite VIII, in males, almost subparallel at sides, rounded­truncate at apex (39 1); internal sac with median sclerites (44 1); rectal loop oblique, straight, provided with a prominent dorsal projection (50 2).

Ptychoderes mixtus differs from the other known species of the genus by the autapomorphies: 1) apical notch of rostrum deep and narrow; 2) elytral interstria 1 prominent from basal 1/5 and continuing with elytral apical margin; 3) setiferous sex patch closer to apical margin of ventrite I. In addition, the whitish dorsal vitta is less contrasting in relation to the body sides.

Ptychoderes mixtus is similar to P. depressus and P. rugicollis in the dorsal vestiture and elytra with anteapical tubercles, differing, in addition to the characters mentioned in the key, in the deep pronotal depression provided with a weakly prominent, almost rounded, median tubercle that does not reach the lateral margins of depression. In P. depressus and P. rugicollis , the pronotal depression is shallower; in the former, the median tubercle is rounded and prominent almost reaching the lateral margins of depression, while in P. rugicollis the median tubercle is transverse and reaches the margins of depression. P. depressus belongs to the P. longicollis + group.

Distribution ( Fig. 266 View FIGURES 265–266. 265 )

PANAMA, COLOMBIA, GUYANA, FRENCH GUIANA, BRAZIL (Mato Grosso, Goiás, Bahia, Espírito Santo, Rio de Janeiro, São Paulo, Paraná, Santa Catarina and Rio Grande do Sul), BOLIVIA, PARAGUAY and ARGENTINA. Jordan (1906) recorded P. mixtus for COSTA RICA and Rheinheimer (2004) for VENEZUELA.

Material examined

PANAMA. Chiriqui Prov., 4 K E Boqueti, 1 F, V .1996, Wappes, Huethei & Morris leg. ( ACMB) . COLOMBIA. 1 M, ex­coll. Lacordaire ( ISNB) . GUYANA. Roraima (distr.), 2 M, ex­coll. Sharp ( BMNH) . FRENCH GUIANA. Cayenne, 1 M, 1 F ( MZSP) . BRAZIL. Mato Grosso: (12°31’S, 51°46’W), 1 M, 1 F, XII GoogleMaps .1968, RS/RGS – Exp. Brazil Beaver leg. ( BMNH); Rio Verde (400 m), 1 M, X.1964, ( DZUP); Rosário Oeste, 1 M, I.1972, Dirings leg. ( MZSP); Vera (12°46’S, 55°36’W), 1 M, 3 F, X.1973, Alvarenga & Roppa leg. ( MNRJ) GoogleMaps . Goiás: Rio Verde , 1 M, II .1945 ( MZSP). Bahia: Encruzilhada (Estr. Rio – Bahia, Km 965, 960 m), 3 M, 2 F, XI .1974, 1 F, XII.1974, Seabra & Roppa leg. ( MNRJ). Espírito Santo: 1 M, ex­coll. Fry ( BMNH); Barra de São Francisco (Córrego Ita), 1 M, XI .1957, Almeida leg. ( DZUP), 1 M, 1 F, XI .1956 , Zikán leg. ( MNRJ); Linhares, 1 F, III – IV .1970 , Silva leg. ( MNRJ); Pinheiros, 1 F, XI .1971 , Lima leg. ( DZUP); Santa Teresa , 1 F, X.1964, 1 F, XII.1964, 1 M, XII.1966, Elias leg. ( DZUP), 1 F, XII .1964 , Elias leg. ( MZSP); São Mateus , 1 M, X.1971, Lima leg. ( DZUP). Rio de Janeiro: 2 M, ex­coll. Fry ( BMNH) . São Paulo: 1 M, I.[19]14, 1 F, I.1920, (leg. illegible) ( MNRJ); Indiana, 1 F, XII .1939, Pohl leg. ( MZSP), 1 M, 1 F, XII .1934 ( MNRJ); Nova Europa (Faz. Itaquerê), 1 M, XII .1964 , Lenko leg. ( MZSP); Piracicaba , 1 M, 1965, Reichardt leg. ( MZSP); São Paulo (Saúde), 1 M, XII . [19]15 ( MNRJ), (Ipiranga) 1 M, ( MZSP), ( Saúde ) 1 m, XII .[19]15 ( MNRJ). Paraná: Antonina, 1 F, I.1956, Marinoni & Azevedo leg. ( DZUP); Curitiba (900 m), 1 M, XII .1967, exp. DZUP – UFPR leg. ( DZUP); Jussara ( Horto Florestal , 340 m), 1 M, X.1974, leg. DZUP UFPR leg. ( DZUP); Ponta Grossa, 1 F (light trap), X.1970, Justus leg. ( DZUP); Porecatu , 1 M, X.1970, Becker­Hatschback leg. ( DZUP); Rolândia, 1 M, 1 F, XII .1943, Justus leg. ( DZUP), 1 M, XI.1949, 1 M, 1 M, III.1951, 2 M, 4 F, VII .1954 , 1 M (compared with syntype), XI.1959, Dirings leg. ( MZSP), 1 M, XII.1957 ( MNRJ). Santa Catarina: Corupá , 2 F, I.1953, 1 F, XI .1953, Maller leg. ( MNRJ); Joinville,1 M, IV .1955, Dirings leg. ( MZSP); Rio Vermelho. 1 M, II .1950, Dirings leg. ( MZSP); Timbó, 4 M, 2 F, II .1956, 2 F, II .1963 , Dirings leg.; Rio Herta (400 m), 1 M, III .1952 , Dirings leg.; Rio Benedito , 2 M, III.1955, Dirings leg. ( MZSP). Rio Grande do Sul: 1 F (compared with syntype), ex­coll. Jordan ( BMNH) . BOLIVIA. Santa Cruz: 4–6 Km SSE Buena Vista (Fauna & Flora Hotel), 1 M, I.2004, Robin & Clarke leg. ( MZSP) . PARAGUAY. Itapúa: E ncarnación, 1 M (compared with syntype), III .1949, Schade leg. ( BMNH); Alto Parana (Hohenau), 1 M, XII .1954, L. Peña leg. ( ACMB). ARGENTINA. Jujuy: Jujuy, 3 M, 2 F, 190[?], Bruch leg. ( MACN), 1 M, 1 F ( DZUP) . Salta: Oran­Urundel, 1 M, XII .1947, Kormilev leg. ( MACN), 1 M ( MZSP).

Ptychoderes bivittatus Jordan, 1894

Figs. 72 View FIGURES 70–84 , 86, 100 View FIGURES 85–112 , 114 View FIGURES 113–126 , 134–137 View FIGURES 127–141 , 220 View FIGURES 215–220 , 228 View FIGURES 228–238 , 248 View FIGURES 239–250 , 265 View FIGURES 265–266. 265

Ptychoderes bivittatus Jordan, 1894: 594; Bovie, 1906: 219 (cat.); Jordan 1906: 304, t.10, fig. 4; Wolfrum, 1929: 6 (cat.); 1953: 3 (cat. supl.); Rheinheimer, 2004: 10 (cat.).

Ptychoderes bivittata [sic]; Blackwelder, 1947: 765 (cat.).

Ptychoderes vittatus [sic]; Jordan, 1895: 123 (misspelling); Wolfrum, 1953: 3 (cat. supl.)

Male. Vestiture ( Figs. 220 View FIGURES 215–220 , 228 View FIGURES 228–238 ): a pale vitta formed by dense, appressed, yellowish scales, intermingled with sparse, semi­appressed light­brown scales present on dorsal side of rostrum, frons, sides of vertex and at each side of pronotum extending to humeri. Central region of vertex and pronotum with brownish scales either moderately dense or missing. E ach elytron with irregular spots formed by short and sub­erect dark­brown scales: at base close to scutellum; one almost rounded antemedially; one dorsal, posmedially, one before apex; several small ones along the pleural margins. Pygidium predominantly with darkbrown scales, except at central region near sulcus, with spot formed by light­brown scales intermingled with scales of other color. Ventral body surface and legs with yellowish scales. Sides of prosternum, below lateral carina, with a continuous vitta of blackishbrown scales, extending shorter beyond lateral carina. Sides of mesosternum with a large glabrous vitta; upper, longitudinal half of metepisternum and lateroanterior angles of ventrites ( Fig. 200 View FIGURES 196–201 ) with dense, decumbent, blackish­brown scales; ventrites, at each side of disk, with shining, brownish scales, forming a row of conspicuous almost rounded spots closer to posterior margin. Mesepimera and mesepisterna with dense, setigerous coarse punctures, each bearing a brownish scale, these punctures shallower on metepisterna, and sides of metasternum and ventrites. Femora, tibiae and tarsi without scales of other color. Outer surface of meso­ and metafemora with two spots of dark­brown scales.

Rostrum 1.3–1.8 times as long as wide across base. Distance between scrobe and eyes from shorter to subequal to maximum width of eye. Sides slightly swollen and bicarinate; upper carina short, lower carina slightly longer. Scrobes careniform at dorsal margin; extension shallow, wide, with two pits, one distal, deep, and one shallower proximal to scrobe. Mentum with moderately coarse, confluent punctures, impressed on each side, shallowly grooved at apical margin. Gula with sparse, fine punctures. Frons without carinae.

Antennae ( Figs. 220 View FIGURES 215–220 , 228 View FIGURES 228–238 ) from short, reaching elytral basal third, to longer than body, exceeding elytral apices by two segments. Segments II–V increasing in length; VI–VII longest, subequal; VIII–IX slightly decreasing in length; IX three times as long as X; this half as long as XI (II = 0.5–1.4; III = 0.7–2.2; IV = 0.6–2.7; V = 0.6–3.1; VI = 0.6–3.5; VII = 0.6–3.5; VIII = 0.6–3.2; IX = 0.8–2.5; X = 0.4–0.7; XI = 0.8–1.6).

Prothorax ( Figs. 220 View FIGURES 215–220 , 228 View FIGURES 228–238 , 248 View FIGURES 239–250 ) as long as or slightly longer than wide; sides subparallel, narrower at anterior margin. Pronotum somewhat convex, without depression, the area corresponding to the median tubercle weakly raised. Pronotum rugose throughout, rugae long, thickened, more conspicuous at middle third; at sides, from feeble to evident. Antebasal carina narrowly interrupted at middle (twice or less than width of carina), oblique towards lateral carina. Lateral carina slightly sinuous. Secondary carina sinuous, narrowly interrupted at middle, almost reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Sides of prosternum, below lateral carina, with irregularly dense, glabrous, shining nodules. Prosternum raised with contiguous, coarse, deep punctures; antecoxal furrow deep, wide, slightly sinuous at middle. Mesosternum with moderately fine, shallow, irregularly sparse punctures, barely impressed on each side, and with two deep, contiguous impressions near intercoxal process. Intercoxal process of mesosternum impressed at apex, slightly swollen at lateral and apical margins, with lateroapical projections. Elytra about twice as long as wide across humeri, somewhat convex, barely depressed along interstria 2; basal gibbosities missing; surface rugose from base to behind middle, more so at base and near suture. Sides subparallel, slightly narrower apically. Apex ( Figs. 220 View FIGURES 215–220 , 228 View FIGURES 228–238 ) with neither anteapical tubercles nor swollen interstriae. Wings without radial cell.

Abdomen. Intercoxal process of ventrite I deeply grooved, sex patch absent. Ventrite V slightly emarginate at apex. Pygidium slightly wider than long, depressed, with an elongate groove to fit elytra; apical margin almost rounded.

Terminalia. Tergite VIII ( Fig. 72 View FIGURES 70–84 ) slightly wider than long, subparallel­sided, roundedtruncate at apex; sternite VIII ( Fig. 86 View FIGURES 85–112 ) almost rounded at sides, slightly emarginate at apex, apodeme triangular, pigmented; apodeme of sternite IX ( Fig. 100 View FIGURES 85–112 ) 3.3 times as long as arms. Tegmen ( Fig. 114 View FIGURES 113–126 ) rounded at apex. Aedeagus ( Figs. 134–137 View FIGURES 127–141 ) with arc between apodemes; tectum ( Fig. 135 View FIGURES 127–141 ) curved at proximal margin, sides acuminate apicad of middle; pedon ( Figs. 134–136 View FIGURES 127–141 ) with body slightly curved, thickened apically, rounded at apex; apodemes slightly curved at proximal half. Internal sac ( Fig. 134 View FIGURES 127–141 ) slightly shorter than aedeagus, spiculated at median and proximal regions; median sclerites thickened, weakly sclerotised ( Fig. 137 View FIGURES 127–141 ).

Female. Rostrum slightly shorter, about 1.2 times as long as wide across base; sides not swollen. Antennae shorter, reaching the antebasal carina of prothorax; segment XI longest (II = 0.4–0.7; III = 0.5–0.8; IV = 0.5–0.9; V = 0.4–0.8; VI = 0.4–0.7; VII = 0.5–0.7; VIII = 0.3–0.6; IX = 0.6–0.9; X = 0.4–0.6; XI = 0.8–1.1). Sides of prosternum, below lateral carina with feeble nodules between pleural suture and posterior margin of prothorax. Ventrite V with apical margin truncate. Terminalia: tergite VIII slightly wider than long, slightly rounded at sides, apical margin feebly emarginated; sternite VIII and ovipositor similar to those of P. mixtus , differing by elongate spermathecal gland, at least twice as long as spermatheca.

Measurements in mm, male/female, respectively. TL = 11.5–19.7/11.5–22.2; RL = 1.7–3.7/1.7–2.9; RAW = 1.7–2.4/1.7–2.5; RBW = 1.3–2.0/1.4–2.2; HW = 1.9–3.0/ 1.9–3.2; DSO = 0.7–1.8/0.6–1.1; MLO = 1.0–1.7/1.0–1.6; PL =3.2–5.5/3.2–5.5; PW = 3.2–5.0/3.2–5.5; EL = 7.5–13.2/7.7–13.5; EW = 3.5–6.0/3.5–6.3.

Type material

Male syntype ( BMNH) examined, with following labels: 1) Type [small white with red border]; 2) Tehuantepec; 3) Ptychoderes bivittatus , male, Type !, Jordan ; 4) [white rounded with yellow border = Paratype]; 5) Karl Jordan coll., B.M. 1940–109; 6) Ptychoderes bivittatus Jordan / Syntype, Lyal det. 1999. Type locality: Jordan (1894) examined a male and a female, respectively from Tehuantepec ( Mexico), and Chontales ( Nicaragua) .

Remarks

P. bivittatus is the sister group of P. callosus + by synapomorphies referent to the anterolateral angles of ventrites I–V with developed, dark spots formed by dense, blackish­brown scales (3 1), and lateral carinae of prothorax strongly sinuous (21 3). It differs from all other species of the genus mainly by dorsal vestiture, frons without carinae, and pronotum slightly convex, lacking central depression.

Distribution ( Fig. 265 View FIGURES 265–266. 265 )

MEXICO, NICARAGUA and COSTA­RICA. Jordan (1906) recorded P. bivittatus for Guatemala and O’Brien (1998) for HONDURAS.

Material examined

CENTRAL AMERICA. 1 F ( NHMB). MEXICO. 2 F, ex­coll. Lange ( NHMB). Colima: Colima, 1 F ( NHMB). Nayarit: Tepic (Elcora), 1 F, Lüdecke leg. ( NHMB). Yucatán: Quintana Roo, 1 F, V.1964 ( ACMB). NICARAGUA: Estelí: Estelí, 1 F, 8/X/2000 (emerged from Ficus insipida – cut by Acrocinus longimanus ), J. Tomá. leg. ( MNRJ); Rio San Juan: Los Guatusos (Rio Papaturro), 1 F, 21–29/ V /2000, Hernandez & Sunyer leg. ( SEAN). COSTA RICA. Guanacaste: Santa Rosa National Park (300 m), 1 M, IV /1984, 1 F, III /1984, ( INBC); 1 F, V /1996 D. H. Janzen & W. Hallwacha leg. ( INBC, MZSP); Est. Biol. Nancite (0 m), 1 M, IX /1990, E. Alcazan leg. ( INBC); P. N. Rincón de la Vieja, Est. Las Pailas (A. C. Guanacaste), 1 M, 8–18 ene/1992, C. Cano leg. ( INBC); Est. Maritza (600 m, west of Vol. Orosi), 1 F, II /1992, F. Araya leg. ( INBC); R. Gongora (700 m, 6 Km NE Quela, Grande de Libéria), 1 M, II /1992, III Curso Parataxon leg. ( INBC, MZSP); P. N. Guanacaste (North of Liberia 30 Km, Finca Jenny), 1 F, II – V /1991, E. Araya leg. ( INBC, MZSP), (240 m) 1 M, 2–6/X/1994, E. Araya leg. ( INBC, MZSP); Vicinity Estac. Murcielago (8 Km SW Cuajiniquil, 100 m), 1 M, VI /1989, GNP Biodiversity Survey UTM leg. ( INBC); Dos de Tilaram (San Ramón, 1.100 m), 1 M, VII /1995, G. Rodríguez leg. ( INBC, MZSP). Limón: Reventazon (“Hamburgfarm”), 1 M, XI.1924, F. Nevermann leg., (“an trockenen Holz”) ( NHMB).

MNHN

Museum National d'Histoire Naturelle

V

Royal British Columbia Museum - Herbarium

MZSP

Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo

DZUP

Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure

MNRJ

Museu Nacional/Universidade Federal de Rio de Janeiro

MACN

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

NHMB

Natural History Museum Bucharest

INBC

Instituto Nacional de Biodiversidad (INBio)

R

Departamento de Geologia, Universidad de Chile

VI

Mykotektet, National Veterinary Institute

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Anthribidae

Genus

Ptychoderes

Loc

Ptychoderes mixtus Jekel, 1855

Mermudes, José Ricardo M. & Napp, Dilma Solange 2006
2006
Loc

Ptychoderes bivittata

Blackwelder, R. E. 1947: 765
1947
Loc

Ptychoderes vittatus

Wolfrum, P. 1953: 3
Jordan, K. 1895: 123
1895
Loc

Ptychoderes bivittatus

Rheinheimer, J. 2004: 10
Wolfrum, P. 1929: 6
Bovie, A. 1906: 219
Jordan, K. 1906: 304
Jordan, K. 1894: 594
1894
Loc

Ptychoderes mixtus

Bovie, A. 1906: 220
Gemminger, M. & Harold, E. 1872: 2722
Jekel, H. 1855: 57
1855