Wadicosa benadira (Caporiacco, 1940) Caporiacco, 1940

Torbjörn Kronestedt, 2015, Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa, European Journal of Taxonomy 132, pp. 1-19 : 3-5

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Wadicosa benadira (Caporiacco, 1940)

comb. nov.

Wadicosa benadira (Caporiacco, 1940) comb. nov.

Figs 1 View Fig. 1 A–C, 2 View Fig. 2 A–D, 3 View Fig. 3 A, D, 4 View Fig. 4 A, D, 5 View Fig. 5 C, D, 7 View Fig. 7 A, 8 View Fig. 8 A, D, 9 View Fig. 9 ;Table 1

Pardosa benadira Caporiacco, 1940: 770 , fig. 1 ( ♂). — Roewer 1959: 60, fig. 18 ( ♂).


The male differs from other Wadicosa species by a short, characteristically shaped embolus with a proximal conical protrusion ( Fig. 2 C – D View Fig. 2 ); the female differs by the shape of the epigyne, having a characteristically shaped median depression which is longer than wide and flanked by two narrow elongate furrows ( Figs 3 View Fig. 3 D, 7 View Fig. 7 A).

Material examined


SOMALIA: ♂, Middle Juba, Jilib (‘Gelib’), 0.49º N, 42.77º E, 20 Apr. 1937 ( MZUF), examined and here designated GoogleMaps .

Other material

KENYA: Isiolo, 5 km NNE of Isiolo, 0º24.3’ N, 37º35.7’ E, 19 Jun. 1999, J.S. Schweikert & W.J. Pulawski, 1 ♀ ( CAS-ENT 9046046 ) GoogleMaps . Laikipia, Mpala Ranch , 0°17‘ N, 37°52’ E, along edge of pond, 20 Apr. 2002, R. Jocqué & C. Warui, 1 ♀ ( MRAC 212184 ) GoogleMaps . Marsabit, Marsabit National Reserve, Lake Paradise , 2°12’ N, 37°56’ E, bare soil on shore, 16 Jan. 1975, T. Kronestedt, 3 ♂♂, 6 ♀♀ (together with Wadicosa oncka ) ( NHRS) GoogleMaps . Kwale, 65 km WNW of Mombasa, at main road to Nairobi , 3°47’ S, 39°17’ E, open grassy vegetation at water hole, 3 & 7 Jan. 1972, T. Kronestedt, 1 ♂, 5 ♀♀ (together with W. oncka ) ( NHRS) GoogleMaps . Taita-Taveta, Tsavo East National Park, Mudanda Rock , 3°10’ S, 38°31’ E, dense grassy vegetation at small pool, 7 Jan. 1972, T. Kronestedt, 2 ♀♀ (together with W. oncka ) ( NHRS) GoogleMaps .

SOMALIA: same data as for lectotype, 1 ♂ (single palp only) GoogleMaps . Middle Shabelle, Garsaale (‘Gaersale’), 2.63º N, 45.37º E, 1 Aug. 1968, S.B.S. (Spedizione Biologica in Somalia del Centro di Studio per la Faunistica ed Ecologia Tropicali del Consiglio Nazionale delle Ricerche) , 1 ♀ ( MRAC 173158 ) GoogleMaps .


Male (Kenya: WNW of Mombasa)

Total length 5.2 mm; carapace 2.70 mm long, 2.10 mm wide.

CEPHALOTHORAX. Carapace brown to blackish brown with yellowish brown, wide, jagged-edged median field. No lateral bands. Sides of thoracic part with short dark recumbent hairs, median field in addition with dense pubescence of adpressed light hairs. Clypeus brown to blackish brown. Chelicerae brown to blackish brown. Sternum yellowish grey.

EYES. Width of row I 50 (slightly procurved as seen from front), row II 62, row III 77, row II–III 59. Diameter of AME 12, ALE 9, PME 24, PLE 19. Distance between AME 9, between AME and ALE 2.

ABDOMEN. Dorsum blackish with pattern of yellowish spots and bars. Lanceolate stripe greyish yellowish. Three yellowish spots at each side of lanceolate stripe, one at anterior half, one at middle and one at rear end. Posterior to lanceolate stripe a series of 3–4 yellowish transverse bars. Venter light yellowish to greyish with adpressed whitish pubescence and more erect scattered light hairs (difficult to see). Spinnerets yellowish.

LEGS (Table I). Yellowish brown with more or less distinct dark annulation on Ti and Mt. Fe I mostly darkened, prolaterally with prominent yellowish spot. (Fe may also be more or less darkened with lighter markings obscured: probably more aged specimens.) Ta yellow. Ti I with two retrolateral spines.

PALP ( Figs 2 C – D View Fig. 2 , 3 View Fig. 3 A, 4 View Fig. 4 A, D, 5 View Fig. 5 C – D). Pt 0.45, Ti 0.45, Cy 1.10. Fe mostly darkened, Pt and Ti more or less darkened yellowish, Cy more or less dark brownish, distally yellowish. Tegular apophysis inconspicuous, largely unsclerotized, widening distally ( Figs 3 View Fig. 3 A, 4 View Fig. 4 A). Anterior retrolateral process comparatively short ( Fig. 4 View Fig. 4 D). Conductor long, narrow, distally bent about 90º ( Fig. 5 View Fig. 5 C – D). Embolus comparatively short, stout, proximally with conspicuous conical protrusion directed backwards, tip with wide concavity ( Figs 2 C – D View Fig. 2 , 5 View Fig. 5 C – D).

Female (Kenya: WNW of Mombasa)

Total length 6.1 mm; carapace 3.25 mm long, 2.35 mm wide.

CEPHALOTHORAX AND ABDOMEN. Carapace greyish brown (slightly lighter than in male). Median field wide, yellowish brown (sometimes weakly discernible), with jagged edges. Lateral bands (when discernible) present as yellowish brown spots, posterior one elongated. Thoracic part richly furnished with short dark hairs; median field and lateral spots in addition with numerous adpressed light hairs. Chelicerae with dark and light hairs of varying length. Colour, pattern and hairiness of abdomen similar to those of male.

EYES. Width of row I 58 (slightly procurved as seen from front), row II 71, row III 89, row II–III 70. Diameter of AME 14, ALE 10, PME 27, PLE 23. Distance between AME 8, between AME and ALE 2.

LEGS (Table 1). Yellowish brown with darker annulation. Fe I mostly dark with distinct yellowish brown spot prolaterally in distal half. Fe II–IV dark except for “pseudoannulation”, i.e., lighter dorso-lateral spots (distally) and dorso-lateral groups of whitish hairs.

EPIGYNE ( Figs 3 View Fig. 3 D, 7 View Fig. 7 A, 8 View Fig. 8 A, D). Median depression longer than wide. Bottom of depression with longitudinal median ridge. Elongated deeper pit at each side of median depression. Lateral elevations with more or less sclerotized rims, protruding like two lips slightly converging rearwards. Anterior part of median depression with two well separated deep foveolae. Receptacula pear-shaped, directed forwards ( Fig. 8 View Fig. 8 D).

Size variation

Carapace lengths of material measured: males 2.65–2.95 mm ( N = 4), females 2.85–3.45 mm ( N = 16).


Collected in habitats close to water.


Somalia, Kenya. ( Roewer (1959) erroneously placed ‘Gelib’ in Ethiopia.)


Italy, Firenze, Museo Zoologico "La Specola"


Sweden, Stockholm, Naturhistoriska riksmuseet


USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum














Wadicosa benadira (Caporiacco, 1940)

Torbjörn Kronestedt 2015

Pardosa benadira

Caporiacco 1940: 770