Scolytodes horridus Jordal and Smith

Jordal, Bjarte H. & Smith, Sarah M., 2020, Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences, Zootaxa 4813 (1), pp. 1-67: 36

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Scolytodes horridus Jordal and Smith

sp. nov.

Scolytodes horridus Jordal and Smith   , sp. nov.

( Figs 57, 60, 63 View FIGURES 55–63 )

Type material. Holotype, female: Ecuador: Los Ríos, Canton Valencia , Reserva Murucumba, S 00°38.544’ W 79°08.902’, 731m, 16.V.2015, Cognato, Smith, Osborn, Martinez et al., ex twigs of Castilla elastica     . Allotype: same label as holotype. Paratypes: same data as HT (7). Holotype, allotype and 1 PT in QCAZ   , 2 PTs in USNM, 2 in ZMBN, and 2 in MSUC.

Diagnosis. Interstriae 10 elevated to near apex of elytra. Protibiae without an additional mesal tooth. Pronotum asperate on anterior half. Elytra with erect interstrial bristles and fine strial and interstrial setae. Distinguished from S. subcribrosus (Eggers)   by the flattened and less setose female frons and the more setose pronotum, and from S. perplexus (Schedl)   , by the asperate pronotum and less setose female frons.

Description female. Length 1.6–1.7 mm, 2.2–2.3 × as long as wide; colour brown. Head. Eyes entire, separated above by 1.9–2.0 × their width. Frons lightly flattened, surface shiny, with moderately coarse punctures from epistoma to upper level of eyes, central half impunctate, glabrous; vestiture confined to scattered, long setae in punctate area. Antennal club with segment 1 corneous, two procurved sutures marked by dense setae. Funiculus 6-segmented. Pronotum reticulated, dull, with obscure punctures on posterior one-third, replaced on anterior twothirds by rough asperities. Vestiture consisting of fine long setae on entire pronotum, and approximately 8 longer, more erect setae (4-2-2). Elytra smooth, shiny; striae regular, not impressed, punctures shallow, spaced by their diameter; interstrial punctures of two sizes, largely in irregular rows, but small punctures confused and contiguous with strial punctures; interstriae 10 elevated to near apex of elytra. Vestiture consisting of fine, erect interstrial setae which is slightly spatulate at the tips, spaced on average by a little less than their length, and by densely placed, irregular rows of fine, semirecumbent, strial and interstrial setae. Legs. Procoxae separated by 0.7–0.8 × the width of one procoxa. Mesocoxae separated by 0.9–1.0 × the width of a procoxa. Protibiae narrow, distal tooth 1 and 2 of equal size, with 3–4 small granules or rugae along the edge towards tibial base; protibial mucro curved posteriorly. Meso- and metatibiae with 5–6 lateral socketed teeth on distal half and third, respectively. Ventral vestiture. Setae on anterior part of metasternum, on mes- and metanepisternum bi-or trifid, on posterior part of metasternum simple.

Male. Similar to female except frons simple, with scant, short setae, surface strongly reticulated.

Key ( Wood 2007). Keys to couplet 48, S. nitellus (Schedl, 1954)   , but the new species is not glabrous and has much more extensive asperities on pronotum.

Etymology. The Latin name horridus   is a masculine nominative adjective, meaning rough or bristly, referring to the rough pronotum with many asperities, and the densely placed erect, subspatulate, interstrial setae on the elytra.

Biology and distribution. Only known from the type locality at median altitude in Ecuador. The holotype was taken from bark of a Castilla elastica   tree ( Moraceae   ).


Museo de Zoologia, Pontificia Universidad Catolica del Ecuador


Smithsonian Institution, National Museum of Natural History


Museum of Zoology at the University of Bergen, Invertebrate Collection