Ooctonus hemipterus Haliday, 1833

Triapitsyn, Serguei V., 2010, Revision of the Palaearctic species and review of the Oriental species of Ooctonus (Hymenoptera: Mymaridae), with notes on extralimital taxa 2381, Zootaxa 2381 (1), pp. 1-74 : 15-20

Ooctonus hemipterus Haliday, 1833

(Figs 9–20)

Ooctonus hemipterus Haliday 1833: 344 ; lectotype female [NMID], designated by Hincks, 1952: 158, not examined (but photographs of the entire brachypterous specimen were examined); type locality: not specified by Haliday (1833), most likely either in England or Ireland.

Eutriche amoena Foerster 1841: 45 ; male [type status not indicated, lost from NHMW], not examined; type locality: Austria, syn. n. Following the opinions of Foerster (1847) and Graham (1982) and based on the short original description of Foerster (1841), I formally synonymize it under O. hemipterus ; E. amoena was listed under O. vulgatus by Dalla Torre (1898).

Ooctonus hemipterus Haliday : Walker 1846: 50 (brief diagnosis, distribution, possible variety of O. vulgatus ); Foerster 1847: 201–202 (brachypterous individuals only); Dalla Torre 1898: 430 (catalog); Debauche 1948: 117, 119 (brachypterous females only); Soyka 1949: 54–55 (brachypterous individuals only); Hincks 1952: 158 (type information, brachypterous individuals only); Hellén 1974: 8 (diagnosis, distribution including a locality in Russia); Graham 1982: 226–227 (also listed Eutriche amoena Foerster as a possible synonym of O. hemipterus ); Donev 1983: 32 (also as O. soykai Hincks ); Viggiani & Jesu 1988: 1024 (records from Italy); Pricop 2008: 44 (illustration), 46 (distribution, taxonomic notes).

Ooctonus vulgatus Haliday : Foerster 1847: 200–201 (misidentification; also supposed synonymy of Eutriche amoena under O. vulgatus ); Soyka 1941: 81–82 (misidentification); Debauche 1948: 119–121 (misidentification); Soyka 1949: 35–36 (figures), 77–78 (misidentification).

Ooctonus atroclavatus Kieffer 1913: 2 ; lectotype female [BMNH], designated by Graham 1982: 227, not examined; type locality:?Clober, Scotland, UK. Synonymized under O. hemipterus by Hincks 1952: 158.

Ooctonus foersteri Soyka 1941: 95–96 [as försteri, invalid spelling]; 2 syntype females [NHMW], examined; type locality: Aachen , North Rhine-Westphalia, Germany, syn. n.

Ooctonus wagneri Soyka 1941: 96 ; holotype female [NHMW], examined; type locality: Sołtysowice (near Wrocław),

Lower Silesian Voivodeship, Poland [indicated by Soyka (1941) as Schottwitz, Schlesien ( Germany)], syn. n. Ooctonus hemipterus f. [forma] hemipterus Haliday : Debauche 1948: 119 (brachypterous female only). Ooctonus hemipterus f. igneus Debauche 1948: 117–118; brachypterous holotype female [INSB], not examined; type locality: Forêt de Loverval, Hainaut, Belgium, syn. n. Ooctonus pechlaneri Soyka 1949: 66–67 ; holotype male [NHMW], examined; type locality: Trins Padaster [South side,

2000 m], Gschnitztal, Tirol, Austria, syn. n. Ooctonus foersteri Soyka : Soyka 1949: 53–54. Ooctonus wagneri Soyka : Soyka 1949: 78–79. Ooctonus soykai Hincks 1952: 160–161 ; holotype female [NHMW], examined; type locality: Valkenburg, Limburg,

Netherlands. Synonymized under O. hemipterus by Graham 1982: 226. Ooctonus amoenus (Förster) : Noyes 2003.

Type material examined. Ooctonus foersteri Soyka : Lectotype female [ NHMW], here designated to avoid confusion about the identity of this species, on slide labeled: 1. “ Ooctonus ( stenomacrus ) 1, 2 (Förster) ♀ foersteri (Soyka) ”; 2. “Type 1”; 3. “Collect. lg. Mayr Förster, Type Aachen, Förster (Canadabals.)”. Paralectotype female [ NHMW] on slide labeled: 1. “ Ooctonus ( stenomacrus ) 1, 2 (Förster) ♀ foersteri (Soyka) ”; 2. “Type 2”; 3. “Collect. lg. Mayr Förster, Type Aachen, Förster det. W. Soyka (Canadabals.)”.

Ooctonus pechlaneri Soyka : Holotype male [ NHMW] on slide labeled: 1. “ Ooctonus pechlaneri (Soyka) ♂ Type”; 2. “Type”; 3. “Gschnitztal, Tirol, Trins Padaster, Südseite, 2000 m, 16.9.1948 lg. Pechlaner in Canadab.”. Note that Soyka (1949) indicated the elevation of the collecting locality of this specimen incorrectly as 2900 m.

Ooctonus soykai Hincks (types designated by Hincks 1952: 160 from the “Genotype”, “Type”, and “Cotype” list for O. vulgatus by Soyka 1941: 82, without labeling them as such, so I labeled them accordingly): Holotype female [ NHMW] on slide labeled: 1. “ Ooctonus vulgatus Haliday “; 2.”Type”; 3. “Valkenburg, Ign. Kolleg, am Fenster, 28. Juni 1932, W. Soyka”. Paratypes on slides: GERMANY. NORTH RHINE- WESTPHALIA: Aachen, G. Mayr: 6 ♂ [ NHMW] 2 (labeled by A. Foerster as “ O. stenomacrus ” 1; 1 ♀ [ NHMW] 3.?Aachen area, G. Mayr: 3 ♀ [ NHMW] 3; 3 ♂ [ NHMW] 2. RHINELAND-PALATINATE, Boppard, G. Mayr, 7 ♂ [ NHMW] 2 (labeled by A. Foerster as “ O. stenomacrus ” 1). NETHERLANDS. LIMBURG, Valkenburg: 18.vi.1931, W. Soyka [5 ♀, NHMW] 2; 10.vii.1931, W. Soyka [1 ♀, NHMW] 2; 28.vi.1932, W. Soyka [1 ♀, NHMW] 2. POLAND. LOWER SILESIA: Krkonoše Mountains [labeled as Riesengebirge (Schlesien)], ca. 1400 m, 27.ix.1933, J. Stammer [1 ♂, NHMW] 2. Małkowice (near Wrocław) [labeled as Malkwitz bei Breslau (Schlesien)], v.1934, J. Stammer [1 ♂, NHMW] 2. OPOLE, Prudnik [labeled as Neustadt (Oberschlesien)], v.1934, W. Soyka [1 ♀, NHMW] 2.

Ooctonus wagneri Soyka : Holotype female [ NHMW] on slide labeled: 1. “ Ooctonus wagneri (Soyka) ♀ ”; 2. “Type”; 3. “Schottwitz – Wiesen b. Breslau, Mai 1934 W. Soyka Canada Balsam.”.

Material examined. AUSTRIA. BURGENLAND, Sankt Andrä am Zicksee , 11.viii.1942, S. Novicky [1 ♂, NHMW] 2 . LOWER AUSTRIA: Hunsdsheim : 2.ix.1941, S. Novicky [1 ♀, USNM] 2 (incorrectly labeled by W. Soyka as “Para-Type”); 10.vii.1961, W. Soyka [1 ♀, NHMW]. Wolfsthal, Rehfeld, 27.ix.1941, W. Soyka [2 ♀, NHMW] 2 . TIROL, Gschnitztal, Trins Padaster ( South side), 2000 m, 16.ix.1948, E. Pechlaner [2 ♀, 1 ♂, NHMW] (one of the females 2; male invalidly labeled by W. Soyka as “co-type” of O. pechlaneri ). BEL- GIUM. LIÈGE: Mont Rigi: 29.vii–4.viii.1983 [1 ♂, CNCI]; 5–11.viii.1983 [1 ♀, 1 ♂, CNCI]; 18–24.viii.1983 [2 ♂, CNCI]; 2–8.ix.1983 [1 ♀, CNCI]. Wanze, Antheit, Corphalie, R. Detry: 14–28.vii.1989 [1 ♂, ISNB]; 28.vii–11.viii.1989 [1 ♀, ISNB]. LUXEMBOURG: Buzenol, 8–24.ix.1981, P. Grootaert [1 ♂, CNCI]. Éthe, Biological Station Bois du Bar, 4.ix.1980, P. Grootaert [2 ♂, CNCI]. Near Éthe and Buzenol, 19.v–16.vi.1981 ,

1. A. Foerster’s manuscript name

2. misidentified by W. Soyka as O. vulgatus Haliday

3. misidentified by A. Foerster as O. vulgatus

P. Grootaert [2 ♀, CNCI]. Ruette, 18–21.v.1981, P. Grootaert [1 ♀, CNCI]. Torgny, 18–21.v.1981, P. Grootaert [1 ♀, CNCI]. CZECH REPUBLIC. CENTRAL BOHEMIA: Calakonce, 30.vii.1991, J. Macek [1 ♂, CNCI]. Mesecin, 8.viii.1992, J. Macek [1 ♂, CNCI]. FRANCE. GIRONDE, Sainte Colombe, 44°54'N 00°02'W, 2.vii.1998, M. van Helden [1 ♀, UCRC]. HERAULT, Montpellier, J.T. Huber: 19–25.iv.1980 [1 ♀, CNCI]; 5– 10.v.1980 [1 ♀, CNCI]. GERMANY. BERLIN, Spandau, viii.1898, T. Pergande [1 ♀, USNM]. LOWER SAXONY, Siedlung Leinetal, Reinshof, ca. 155 m, 17.vi.1953, Prilop (in sugar beet field) [1 ♀, NHMW]. NORTH RHINE-WESTPHALIA: Aachen, G. Mayr: 1 ♀ [ NHMW] (det. A. Foerster); 1 ♀ [ NHMW] (det. W. Soyka).? Aachen area, G. Mayr: 4 ♀ [ NHMW] (det. W. Soyka); 4 ♀ [ NHMW] (det. A. Foerster). Burscheid, viii.1959, M. Boness [1 ♂, NHMW]. Refrath (near Bensberg), M. Boness: 25.v.1955 [1 ♀, NHMW]; 25.v.1960 [1 ♀, 1 ♂, NHMW]. RHINELAND-PALATINATE: Boppard, G. Mayr [1 ♂, NHMW] (labeled by A. Foerster as “ O. angustipennis ” 1 and identified by W. Soyka as O.? pechlaneri ). Near Mainz, W. Steffan [1 ♀, CNCI]. SCHLESWIG-HOLSTEIN, Schödlbach (near Kiel), viii.1954, M. Boness [1 ♀, NHMW]. No location indicated (presumed in Germany), 2 ♂ on minuten pins [ MHNG] (labeled by A. Foerster as “ O. stenomacrus ” 1). GREECE. CENTRAL MACEDONIA, Lake Kerkini, G. Ramel: Beles Mts. , 41°17’19.5’’N 23°12’18.4’’E, 550 m, 9–15.v.2005 [1 ♂, UCRC]; Ecotourism site, 41°08’15.6’’N 23°13’01.2’’E, 65 m: 2– 8.v.2006 [3 ♀, 1 ♂, UCRC]; 9–15.v.2006 [5 ♂, BMNH, UCRC]; 16–22.v.2006 [1 ♂, UCRC]; 13.v–5.vi.2006 [2 ♂, UCRC]; Kerkini Marsh, 41°13’32.8’’N 23°05’04.2’’E, 45 m: 11–17.iv.2007 [2 ♀, UCRC]; 18– 24.iv.2007 [8 ♀, BMNH, UCRC]. HUNGARY. VAS, near Köszeg, 16–19.v.2001, L. Masner [1 ♀, CNCI]. ITALY. CAMPANIA, CASERTA, SE end of Lago Matese, 41°24.41'N 14°24.20'E, 1050 m, 7–8.vi.2003, M. Bologna, J. Munro, A. Owen, J.D. Pinto [3 ♂, UCRC]. PIEDMONT, NOVARA, Lago d'Orta, 0.5 km S of Invorio-Inferiore, 410 m, 5.x.1991, H. Baur [1 ♂, CNCI]. UMBRIA, PERUGIA, Monte Puglia, 8–23.vi.1978 [1 ♂, CNCI]. KYRGYZSTAN. DZHALAL-ABAD, Beke-Chal, 41°32'50''N 72°29'38''E, 960 m, 23.vi.1999, C.H. Dietrich [1 ♂, UCRC]. ISSYK-KUL, Terskey Alatau, Turuk-2 Pass, 42°24'35''N 79°24'30''E, 3370 m, 20.vii.2000, C.H. Dietrich [3 ♂, UCRC]. NARYN: at Alabuga River , 25 km W of Baetovo, 41°17'47''N 74°39'20''E, 1700 m, 16.vii.2000, C.H. Dietrich [2 ♂, UCRC]. Dzhumgal-Too Ridge, Seok River (E ravine), 42°13'13''N 75°00'55''E, 2620 m, 25.vii.2000, C.H. Dietrich [1 ♀, UCRC]. OSH, N slope of Trans Alaj Ridge near Bordobo, 39°29'56''N 73°16'30''E, 3520–3743 m, 11.vii.2000, C.H. Dietrich [1 ♀, UCRC]. MOROCCO. MARRAKECH-TENSIFT-EL HAOUZ, Ouirgane, 31°08'N 08°05'W, 1000 m, 1–7.ii.1997, C. Kassebeer [1 ♂, CNCI]. NETHERLANDS. GELDERLAND, Wageningen, 26.vii.1978, L. Huggert [2 ♂, CNCI]. LIM- BURG, Valkenburg, vi.1932, W. Soyka [2 ♀, EMEC, NHMW] 2. UTRECHT, Leersum, Brockhuizen, viii.1975, H.J. Vlug [1 ♀, CNCI]. POLAND. Boczan [sic, province unknown], 23.vii.1927, Schulezewsky [1 ♂, NHMW] 2. ŁÓDŻ, Skierniewice, 1920s, S. Novicky [1 ♀, 1 ♂ 2, EMEC; 1 ♀, 3 ♂ 2, NHMW]. MASOVIA: Grójec, 21.vi.1935, S. Novicky [1 ♀, NHMW] 2. Warsaw, S. Nowicky: 1933 [1 ♀, NHMW]; viii.1935 [1 ♀, 2 ♂, NHMW] 2. RUSSIA. LENINGRADSKAYA OBLAST’: Pushkin, 14.ix.1985, E.S. Sugonjaev, J. LaSalle [2 ♂, CNCI]. 69-km [Railway Station] near Sosnovo, 25–26.viii.1985, V.A. Trjapitzin [1 ♂, ZIN]. MOSKOVS- KAYA OBLAST’: NOGINSKIY RAYON, Fryazevo, M.E. Tretiakov: 2–15.vi.2000 [8 ♀, UCRC]; 25.vi– 2.vii.2000 [1 ♀, 1 ♂, UCRC]; 7–15.vii.2000 [2 ♀, UCRC]; 24.vii.2000 [1 ♂, UCRC]; 26.vii–14.viii.2000 [1 ♀, UCRC]; 15–25.viii.2000 [3 ♀, UCRC]; 25–31.viii.2000 [2 ♀, UCRC]; 21.vi.2001 [2 ♀, UCRC]; 20.vii.2001 [1 ♀, UCRC]; 24.viii.2001 [1 ♀, UCRC]. PUSHKINSKIY RAYON, Mamontovka, E.Ya. Shouvakhina: 10–20.vii.2000 [2 ♀, UCRC]; 10–20.viii.2000 [1 ♀, UCRC]; 20–31.viii.2000 [1 ♀, ZIN]; 26.v– 5.vi.2001 [1 ♀, 1 ♂, ZIN]; 15–25.vii.2001 [1 ♀, UCRC]. PRIMORSKIY KRAY, Gornotayozhnoye (18 km SE of Ussuriysk), 43.66°N 132.25°E, 200 m, M.V. Michailovskaya: 11–12.vi.2000 [1 ♀, 1 ♂, UCRC]; 18– 19.vi.2000 [1 ♀, UCRC]; 2–4.x.2000 [1 ♂, UCRC]. SAKHALINSKAYA OBLAST’, Kuril Islands : Rasshua Island , near (inland of) Malen’kaya Bay , 47°42.77’N 152°58.16’E, 4.viii.1999, B. Urbain [1 ♂, CAS]. Shiashkotan Island , SW (inland of) Zakatnaya Bay , 48°46.77’N 154°01.99’E, 29.vii.2000, T.R. Anderson [1 ♂, CAS]. SLOVENIA. UPPER CARNIOLA, Kamna Gorica, 1–13.viii.1978, L. Huggert [1 ♂, CNCI]. SWE- DEN. UPPSALA, Eriksberg, 11–19.vii.1987, F. Ronquist [1 ♂, CNCI]. VÄRMLAND, Ekshärad, 17 –

22.vii.1960, W.R.M. Mason [2 ♀, CNCI]. SWITZERLAND. BERNE: Berne (Bern), 11.vi.1924, C. Ferrière [1 ♀, NHMW] 2. Kandersteg, 1790 m, 29.viii.1991, H. Baur [1 ♂, CNCI]. Lenk, Buhlberg, 1800 m, 14.viii.1981, R. Detry [1 ♀, CNCI]. ZÜRICH, Dielsdorf, 650 m, 17.viii.1984, L. Masner [1 ♂, CNCI]. TUR- KEY. ANTALYA: Kemer, 14–20.v.1991, J. LaSalle [1 ♀, 2 ♂, CNCI]. Phaselis, 16.v.1991, J. LaSalle [1 ♂, CNCI]. UNITED KINGDOM. ENGLAND: BERKSHIRE, Ascot , Silwood Park , 13.viii.1969, Dietrick (“ Holcus patch”) [3 ♀, 2 ♂, BMNH]. BUCKINGHAMSHIRE, Burnham Beeches , 12.vi.1914, C.O. Waterhouse [2 ♂, BMNH]. CAMBRIDGESHIRE, Abbots Ripton , Monks Wood National Nature Reserve , 17– 28.vii.1978, M. Fitton, J.S. Noyes [1 ♀, BMNH]. DORSET: Bournemouth, S.G.C. Brown: 12.ix.1979 [1 ♀, BMNH]; viii.1981 [2 ♂, BMNH]; 8.x.1981 [2 ♀, BMNH]; 1982 [1 ♀, BMNH]. West Woods , 21.vi.1952, S.G.C. Brown [1 ♀, BMNH]. HAMPSHIRE, Brockenhurst , 26.v.1976, J.S. Noyes [1 ♂, BMNH]. SURREY: Box Hill (near Dorking ): 4.ix.1988, S.L. Heydon [1 ♂, UCDC]; 4.ix.1988, J.S. Noyes [1 ♀, UCDC, 2 ♂, CNCI, UCRC]; 16.vii.1994, J.S. Noyes [1 ♂, CNCI]. Richmond, C.O. Waterhouse: 5.ix.1908 [1 ♀, BMNH]; 7.x.1910 [1 ♂, BMNH]. WEST SUSSEX, Worthing , 7.x.1910, C.O. Waterhouse [1 ♂, BMNH]. No locality or any other data indicated [presumably from England] [1 ♂, BMNH] (slide-mounted and misidentified as O. vulgatus by F. Enock). WALES: CLWYD: Dolywern (5 km S of Llangollen), 29.vii.1999, J.S. Noyes [1 ♂, CNCI]. Lamon Dyffryn Ceiriog (10 km SW of Llangollen), 31.vii.1999, J.S. Noyes [1 ♂, CNCI]. DENBIGH, Abergele Coed y Gopa , 27.vii.1999, J.S. Noyes [2 ♂, CNCI]. GLAMORGAN: Kenfig Pool National Nature Reserve , J.S. Noyes: 6.viii.1988 [3 ♀, 3 ♂, CNCI]; 4.viii.1994 [1 ♀, 1 ♂, UCRC]; 18.viii.1998 [4 ♂, CNCI]; 31.viii.1998 [1 ♂, CNCI]. Oxwich National Nature Reserve , 5.viii.1994, J.S. Noyes [3 ♂, CNCI]. Whiteford Burrows National Nature Reserve , 2.viii.1988, J.S. Noyes [5 ♂, CNCI (3), UCRC (2)]. POWYS, 10 km W of Brecon, 15.ix.1988, S.L. Heydon [3 ♂, UCDC] .

Extralimital records. CANADA. NORTHWEST TERRITORIES, at Horton River , 68.00°N 123.28° W, 18–20.vii.2000, B.V. Brown [1 ♂, UCRC] . USA. ILLINOIS, KANE, 1 mi. S of Algonquin at Fox River , 18.vi.1991, H.E. Andersen [1 ♂, UCRC] .

Redescription. FEMALE (lectotype and paralectotype of O. foersteri , and non-type specimens). Body length 900–1150 µm. Head and mesosoma very dark brown or black except eyes and ocelli pale or dirty pink; petiole yellowish or yellowish brown; gaster brown to dark brown; radicle light brown, the rest of scape and pedicel brown, and flagellum dark brown; legs yellowish brown except distal tarsomeres brown.

Head with more or less defined reticulate sculpture.

Antenna (Fig. 9) shorter than body, with radicle 0.2–0.26x length of scape, remainder of scape 4.1–5.6x as long as wide, a little wider medially than basally or apically, longitudinally striate; pedicel much longer than each of F1–F6 and a little longer than F7 or F8; all funicle segments longer than wide, F1–F6 more or less subequal in length and without mps, F7 and F8 notably longer and wider than preceding funicle segments, each with 2 mps; clava pointed in dorsal view and oval in lateral view, 2.4–3.0x as long as wide, shorter than scape plus radicle, about as long or slightly shorter than combined length of F5–F8 and about as long or slightly longer than combined length of F1–F5, with 7 mps.

Entire mesosoma (Fig. 10) with strong reticulate sculpture; propodeum (Fig. 11) without median carina, the median areole thus very large, narrowing and extending to anterior margin of propodeum, lateral carinae subparallel, extending to anterior margin of propodeum.

In brachypterous specimens, wings variably abbreviated, extending from just beyond mesosoma to about two thirds length of gaster according to Hincks (1952) [in the lectotype, forewing extending to about two thirds length of gaster]; in most brachypterous specimens examined, forewing extending to about one third length of gaster, with disc reduced just beyond apex of venation and with a strong brownish tinge (Fig. 12); hind wing with disc reduced at apex of venation and with a strong brownish tinge (Fig. 13). In macropterous specimens, forewing ( Fig. 14 ) extending beyond apex of gaster, 3.3–3.5x as long as wide; disc with a light to strong brownish tinge throughout, densely setose except bare behind base of submarginal vein, rounded apically; longest marginal seta 0.26–0.37x greatest width of wing. Hind wing ( Fig. 14 ) 17–18x as long as wide;

disc with a slight brownish tinge and setose throughout except less setose behind apex of venation; longest marginal seta 2.2–2.4x greatest width of wing.

FIGURES 9–13. Ooctonus hemipterus ♀ (9, 10, 12, 13 – 39°29’56’’N 73°16’30’’E, Osh, Kyrgyzstan; 11 – Kenfig Pool National Nature Reserve, Wales): 9, antenna; 10, mesosoma; 11, posterior scutellum, metanotum, and propodeum; 12, forewing; 13, hind wing.

Coxae and metafemur reticulate, pro- and mesofemora longitudinally striate.

Gaster ( Fig. 15 ) at least a little longer than mesosoma; petiole ( Fig. 15 ) lightly to notably sculptured, 2.3– 3.1x as long as wide, 1.0–1.2x as long as metacoxa; ovipositor occupying 0.8–0.9x length of gaster, barely exserted beyond apex of gaster (by less then 0.1x own length), 1.1–1.3x length of metatibia.

MALE (non-type specimens). Body length 850–1100 µm. Similar to female except for the normal sexually dimorphic features and the following. Antenna ( Fig. 16 ) longer than body, with scape plus radicle 4.9– 5.3x as long as wide, scape and pedicel brown, flagellum dark brown; mesosoma as in Fig. 17 ; only macropterous forms known, forewing ( Fig. 19 ) 3.7–4.4x as long as wide; hind wing as in Fig. 20 ; legs often partially a little darker than in female; gaster shorter than in female; genitalia as in Fig. 18 .

Diagnosis. Ooctonus hemipterus is characterized by the reticulate head and mesosoma (including the propodeum) and presence of brachypterous (females only) or macropterous (both sexes) forms; a normal (non-abbreviated) forewing has a rounded apex ( Fig. 14 ).

Distribution. PALAEARCTIC: Austria, Belgium, Czech Republic, France *, Germany, Greece *, Hungary *, Italy, Kyrgyzstan *, Morocco *, Netherlands, Poland, Russia, Slovenia *, Sweden, Switzerland *, Turkey *, UK ( England, Scotland, and Wales *); unconfirmed records (no material has been examined): Bulgaria ( Donev 1983), Ireland ( Walker 1846), Croatia, Northern Ireland ( UK), Slovakia ( Noyes 2003), Finland ( Hellén 1974), and Romania ( Pricop 2008). NEARCTIC*: Canada *, USA *.

Hosts. Unknown.

Figures

FIGURE 14. Ooctonus hemipterus ♀ (Mamontovka, Moskovskaya oblast’, Russia): wings.

FIGURE 15. Ooctonus hemipterus ♀ (Fryazevo, Moskovskaya oblast’, Russia): metasoma (scanning electron micrograph).

FIGURES 16–20. Ooctonus hemipterus ♂ (16, 17 – Oxwich National Nature Reserve, Wales; 18–20 – Kenfig Pool National Nature Reserve, Wales): 16, antenna; 17, mesosoma; 18, genitalia; 19, forewing; 20, hind wing.