Thomasomiris, Minghetti & Montemayor & Dellapé, 2024
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publication ID |
https://dx.doi.org/10.3897/dez.71.104130 |
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publication LSID |
lsid:zoobank.org:pub:A0C8103C-B56C-40E9-9C38-1D2617F8A3C4 |
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persistent identifier |
https://treatment.plazi.org/id/9FF34EA5-DAB6-426D-87B7-D9BD97CA0EC5 |
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taxon LSID |
lsid:zoobank.org:act:9FF34EA5-DAB6-426D-87B7-D9BD97CA0EC5 |
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treatment provided by |
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scientific name |
Thomasomiris |
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gen. nov. |
Thomasomiris gen. nov.
Type species.
Thomasomiris setosus sp. nov.
Diagnosis.
Head, collar, posterior pronotal lobe, scutellum, and hemelytra with short, erect setae; eyes small, stylate, directed anteriorly (Fig. 1A View Figure 1 ), less than half head height (Figs 2A View Figure 2 , 3A View Figure 3 ); vertex with a medial longitudinal depression; antennal segment I about half the length of vertex width and almost 4 times as short as antennal segment II; anterior pronotal lobe depressed in lateral view; cuneal fracture extending to middle of corium; cuneus wider than long (Fig. 1A View Figure 1 ); veins angled at the middle of membrane, slightly surpassing apex of cuneus; and tibiae wide, with abundant semierect setae.
Description.
Macropterous female. Body length to apex of membrane 5.6-5.7; body length to apex of cuneus 4.6-4.7.
Surface shiny, pilose; posterior pronotal lobe evenly punctate. Head with short and erect setae (Fig. 3A View Figure 3 ); antennal segment I with recumbent setae on distal half (Fig. 3A View Figure 3 ), antennal segments II-IV with short, recumbent and longer, semierect setae. Collar, posterior pronotal lobe, scutellum and hemelytra with short, erect setae.
Structure. Head width more than twice its length. Clypeus flat in lateral view (Fig. 3A View Figure 3 ), not visible from above. Frons flat in lateral view. Eyes small, less than half head height (Figs 2A View Figure 2 , 3A View Figure 3 ), stylate, directed upwards and anteriorly, interior and posterior margins straight, interior margin at same level with lateral collar margins (Fig. 1A View Figure 1 ). Maxillary plates rectangular, large, long. Labium extending to base of abdomen. Antennal segment I shortest, about half the length of vertex width, narrower basally; segment II almost 4 times as long as antennal segment I; segment III thinner than II (Fig. 1A View Figure 1 ); segment IV shorter and thinner than III. Thorax: pronotum trapeziform, slightly directed downwards. Anterior pronotal lobe clearly distinguished from posterior pronotal lobe, depressed in lateral view (Fig. 3A View Figure 3 ); collar wider than antennal segment I; calli evident, medium-sized, depressed and separated from each other by a median depression. Posterior pronotal lobe flat, with shallow lateral depressions before humeral angles, posterior margin convex (Fig. 1A View Figure 1 ). Hemelytron flat in lateral view; lateral margin convex (Fig. 1A View Figure 1 ); embolium flat, thin and slightly expanded in posterior margin; medial fracture visible, adjacent to R+M vein; cuneus wider than long (Fig. 1A View Figure 1 ); veins angled at middle of membrane and directed posteriorly behind posterior margin of cuneus. Legs: femora flat; metafemora slightly curved; tibiae robust; protibiae flat on distal interior face.
Male. Unknown.
Geographic distribution.
Panama (Fig. 11 View Figure 11 ).
Etymology.
The name of the new genus is formed from the given name of our friend and colleague Dr Thomas J. Henry, who has published numerous important papers on Heteroptera , including many on Miridae , combined with miris in reference to its assignment to the family Miridae . Gender masculine.
Discussion.
The first couplet of the key to the Neotropical eccritotarsine genera ( Carvalho and Ferreira 1995) refers to the development of male hemelytra; we assumed that males of Thomasomiris have normal hemelytra, with clavus and corium distinguished and membrane present, so we followed to couplet 3. The two monotypic genera identified in the couplet 2, Aztecariella Carvalho, 1951 and Coleopteromiris Carvalho, 1946 show different characters from Thomasomiris , as a pronotum coarsely punctate and females without division of clavus, corium and cuneus, and without membrane in Aztecariella , and sessile eyes and females with coleopteroid hemelytra in Coleopteromiris . Following the key, Thomasomiris runs to the couplet 8 where the genera Hesperolabops Kirkaldy, 1902 and Aztecarina Calvalho, 1974 are identified and are discriminated by male genitalia characters, but unlike Thomasomiris , Hesperolabops shows strongly pedunculated eyes, a lobate collar, a longer antennal segment I, and a cuneus longer than width, and Aztecarina shows a strongly punctate pronotum with its posterior margin sinuated and antennal segments I and II of similar length. Four Neotropical genera were omitted in the Carvalho and Ferreira’ key, Perissobasis Reuter, 1892, Eurycipitia Reuter, 1905, Bugabacoris Carvalho & China, 1959 and Pycnoderiella Henry, 1993, and five genera were described after 1995, Cubanomiris Hernández & Stonedahl, 1996 and Agaveocoris , Laterospinocoris , Nigrotomocoris and Schaffnerocoris , described by Henry and Menard 2020 including some species with neotropical distribution. The medium size, the stylate eyes, the labium extending at least to the metacoxae, and the convex lateral margins of hemelytra are similar to those found in species of Neoneella Costa Lima, 1942. However, the short, erect setae, the upward and anteriorly directed eyes located far from the collar, the antennal segment II almost 4 times as long as antennal segment I, the flattened anterior pronotal lobe clearly separated from the evenly punctate posterior pronotal lobe with the posterior margin convex, the short, wide cuneus, and the concave posterior margin of cell on membrane forming an acute angle at the middle, distinguish this new genus from Neoneella and from all other eccritotarsines.
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