Anelastes abbreviatus, Kovalev, 2019

Kovalev, Alexey V., 2019, A review of the genus Anelastes Kirby, 1819 (Coleoptera: Eucnemidae) of the Palaearctic fauna, Zootaxa 4683 (1), pp. 97-119 : 100-101

Anelastes abbreviatus sp. nov.

( Figs 1–3 , 15 , 24–26 , 40 , 48, 49 , 57, 58 , 69, 70, 73 , 93, 94, 98 )

Type material. Holotype, ♂: ALGERIA: Tlemcen Province: “ Daya [= Dhaya]” [p] // “Muséum Paris / Coll. L. Bedel 1922” [p, blue label] ( MNHN); paratype, ♀: “Daya [p] / Dr. Munier [hw]” // “Muséum Paris / Coll. L. Bedel 1922” [p, blue label] // “barbarus / Luc.” [hw, blue label] ( MNHN).

Differential diagnosis. Among the Palaearctic congeners this new species is distinguishable due to its incomplete lateral carinae of the pronotum, elytra with odd-numbered discal interstriae somewhat wider than even-numbered ones, widely separated antennal insertions, comparatively short antennae and legs, smaller eyes, as well as robust and strongly convex body with denser granulation of the integument. The another known species with incomplete lateral pronotal carinae, Nearctic A. drurii , differs from the new species in the less convex and more elongate body, longer antennae and legs, sparser granulation of integument, and especially, in shape of aedeagus and structure of female abdominal segment VIII (see Cobos 1965; Chassain 1978; Muona 2000). The aedeagus of A. abbreviatus sp. nov. more resembles those in A. alius sp. nov., A. californicus and A. desertorum , however, the penis in these species are nearly straight posteriorly.

Description of holotype. Body length 6.5 mm. Coloration uniformly ferruginous; body integument dull, with dense, minute decumbent yellowish setae.

Head 0.70 times as wide as pronotum; integument with dense to very dense granulation. Distance between inner edges of antennal insertions 0.5 times as great as that between mandibular bases ( Fig. 15 ). Eyes relatively small, slightly convex ( Figs 15 , 40 ). Antennae short, slightly extending posteriorly beyond the middle of pronotum, moniliform ( Figs 24 ). Antennomere 3 elongate, thickened apically, 1.30 times as long as wide and 1.35 times as long as antennomere 4. Antennomeres 4–8 strongly thickened apically, 0.9–1.0 times as long as wide. Antennomeres 9–10 about as long as wide. Antennomere 11 1.20 times as long as antennomere 10. Orifices of sensory cavities of antennomeres 9–11 strongly transverse, slit-like.

Pronotum transverse, 1.35 times as wide as long at midline, widest behind middle; sides strongly convex and slightly sinuate at posterior angles; disc strongly convex, with more or less impressed median groove in posterior third of pronotum and with weak paramedian impressions. Lateral carinae of pronotum distinctly bent anteriorly and obliterated in posterior half ( Fig. 40 ). Posterior angles short, slightly divergent, rounded at apex. Integument with granules very dense at middle, and becoming denser and more prominent towards sides and apex; interspaces between granules at middle about as wide as granules. Prosternum and hypomera covered with granules dense, interspaces between granules 1.0–1.5 times as wide as granules, granulation of hypomera somewhat finer.

Elytra twice as long as wide combined and 2.75 times as long as pronotum, subparallel-sided and widest about midlength. Elytral striae impressed, with moderately strong, coarse and dense punctures; odd-numbered discal interstriae somewhat wider and more convex than even-numbered ones; integument with very dense granulation, interspaces between granules about as great or less than one granule diameter. Granulation of metaventrite and visible portion of metepisterna somewhat finer, and about as dense as those on prosternum.

Legs comparatively short; tarsi short with tarsomeres strongly widened apically; metatarsus about 0.8 times as long as metatibia ( Figs 48, 49 ).

Abdominal ventrites at middle with granules moderately dense, becoming denser on ventrite 5; interspaces between granules 1–3 times as wide as granules. Ventrite 5 as in Fig. 57 . Tergite VII (pygidium) apically narrowly rounded. Sternite 9 slightly notched at apex. Aedeagus as in Figs 69, 70, 73 .

Female. Body length 9.1 mm. Antennae as in Fig. 25 . Pronotum transverse, 1.35 times as wide as long at midline, with paramedian impressions on pronotal disc nearly indistinct; lateral carinae of pronotum distinctly bent anteriorly and obliterated in posterior third. Elytra 2.05 times as long as wide combined and 2.75 times as long as pronotum, with odd-numbered discal interstriae somewhat wider than even-numbered ones, interstriae equally slightly convex. Ventrite 5 as in Fig 58 . Tergite VIII rounded at apex ( Fig. 94 ). Abdominal segment VIII as in Figs 93, 94 ; sclerites of sternite VIII comparatively small and lightly sclerotized ( Fig. 94 ).

Etymology. The epithet is a Latin adjective meaning “shortened”.

Distribution. Algeria (Tlemcen Province).

Figures

FIGURES 1–3. Anelastes abbreviatus sp. nov. Fig. 1. Holotype, male. Fig. 2. Labels of holotype. Fig. 3. Paratype, female. Scale bar = 2.0 mm.

FIGURES 15–18. Anelastes spp., head, frontal. Fig. 15. A. abbreviatus sp. nov., holotype, male. Fig. 16. A. alius sp. nov., holotype, male. Fig. 17. A. barbarus, male from Maroc (Oued Iquem). Fig. 18. A. crenulatus, male from Cambodia (“Phnum Basset”). Scale bars = 1.0 mm.

FIGURES 24–28. Anelastes spp. Fig. 24. A. abbreviatus sp. nov., male antenna, holotype. Fig. 25. A. abbreviatus sp. nov., female antennomeres 6–11, paratype. Fig. 26. A. abbreviatus sp. nov., posterior surface of antennomeres 9–11, schematized. Fig. 27. A. alius sp. nov., male antenna, holotype. Fig. 28. A. alius sp. nov., posterior surface of antennomeres 9–11, schematized. Scale bars = 0.5 mm.

FIGURES 40–43. Anelastes spp., prothorax, lateral. Fig. 40. A. abbreviatus sp. nov., holotype, male. Fig. 41. A. alius sp. nov., holotype, male. Fig. 42. A. barbarus, male from Maroc (Oued Iquem). Fig. 43. A. crenulatus, male from Cambodia (“Phnum Basset”). Scale bars = 1.0 mm.

FIGURES 47–55. Anelastes spp. Fig. 47. A. barbarus, hindwing. Fig. 48. A. abbreviatus sp. nov., male metatibia and metatarsus, lateral. Fig. 49. A. abbreviatus sp. nov., male metatarsus, dorsal. Fig. 50. A. alius sp. nov., male metatibia and metatarsus, lateral. Fig. 51. A. alius sp. nov., male metatarsus, dorsal. Fig. 52. A. barbarus, male metatibia and metatarsus, lateral. Fig. 53. A. barbarus, male metatarsus, dorsal. Fig. 54. A. crenulatus, male metatibia and metatarsus, lateral. Fig. 55. A. crenulatus, male metatarsus, dorsal. Scale bars = 1.0 mm.

FIGURES 56–63. Anelastes spp. Fig. 56. A. barbarus, elytral apices and abdominal apex, dorsal. Fig. 57. A. abbreviatus sp. nov., male ventrite 5, holotype. Fig. 58. A. abbreviatus sp. nov., female ventrite 5, paratype. Fig. 59. A. alius sp. nov., male ventrite 5, holotype. Fig. 60. A. barbarus, ventrite 5 of male from Maroc (Oued Iquem). Fig. 61. A. barbarus, ventrite 5 of female from Maroc (Oudjda). Fig. 62. A. crenulatus, ventrite 5 of male from Cambodia (“Phnum Basset”). Fig. 63. A. crenulatus, ventrite 5 of male from Myanmar (“Thagata”). Scale bars = 1.0 mm.

FIGURES 69–74. Anelastes spp., males. Figs 69, 70, 73. A. abbreviatus sp. nov., holotype. Figs 71, 72, 74. A. alius sp. nov., holotype. Figs 69, 71. Aedeagus, lateral. Figs 70, 72. Apex of aedeagus, dorsal. Figs 73, 74. Paramere, lateral. Scale bars = 0.5 mm.

FIGURES 93–98. Anelastes spp., females. Figs 93, 94, 98. A. abbreviatus sp. nov. Figs 95, 97. A. barbarus. Fig. 96. A. crenulatus. Fig. 93. Abdominal segment VIII, ventral. Figs 94–96. Abdominal segment VIII dorsal (left) and ventral (right). Fig. 97. Ovipositor, ventral. Fig. 98. Genital tract. bc—bursa copulatrix; opd—duct leading to ovipositor; ovd—duct leading to ovaries; S8—sternite 8; sg—spermathecal gland; sp—spermatheca; T8—tergite VIII. Scale bars = 0.5 mm.