Goja andina, Carrejo & Obando & Casasola-González & García Aldrete, 2021

Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo & García Aldrete, Alfonso N., 2021, New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico, Zootaxa 5040 (4), pp. 451-481 : 461-463

Goja andina n. sp. Male

( Figs 1–6 )

Diagnosis. With G. risaraldensis n. sp., and G. vallecaucana n. sp., it constitutes an assemblage of related species, characterized by having the external parameres well developed and by having endophallic sclerites unlike those observed in the known species of Goja . It differs from the two species above by having R4+5 of the forewings seven branched; by having the external parameres mid sized, by having the apex of the aedeagal arch with small, pointed projections directed outward, with the posterior border straight, and by having the right endophallic sclerites with two mesal processes, one shorter than the other ( Fig. 6 ).

Color (in 80% ethanol). Body brown, with light spots.

Morphology. Head: H/MxW: 1.64; compound eyes small, H/d: 3.61; IO/MxW: 0.66. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.21. Lower edge of labrum with a pair of projections with rounded apices. Forewings ( Fig. 1 ): FW/W: 2.88; pterostigma elongate, wider in the middle: lp/wp: 6.19, Rs supernumerary with six or more branches, basal portion of Rs concave after transverse r-m; areola postica elongate, may be open and partially fused to M: al/ah: 4.00; M supernumerary. Hindwings ( Fig. 2 ): HW/w: 3.31. Rs five branched; M four branched; Rs and M joined by a crossvein, asymmetry in wing venation may be present. Legs: fore tarsi without ctenidiobothria, mid and hind tarsi with ctenidiobothria. Hypandrium broad, setae as illustrated ( Fig. 5 ) posterior border almost straight. Phallosome ( Fig. 6 ): side struts straight; aedeagal arch robust, apex with two short mesal projections, acuminate, directed outward; external parameres well developed, concave-convex, asymmetric, right one sclerotized, acuminate distally, left one shorter. Three pairs of asymmetric endophallic sclerites ( Fig. 6 ): right pair with two mesal processes, lanceolate, apically acuminate, one shorter than the other; two lateral sclerites, outermost apically wide and projected posteriorly, anteriorly narrow; innermost narrow and distally lobed, base folded on itself as a reticulate lamina; left sclerites mesally lanceolate, apex acuminate; two outermost lateral sclerites wide apically, narrow mesally, anteriorly wide with fine spicules. Clunium projected posteriorly over the area of the epiproct. Paraprocts ( Fig. 4 ), oval, with setae and macrosetae on posterior border, sensory fields large, circular, with 32 trichobothria in basal rosettes. Epiproct slightly concave anteriorly, rounded posteriorly, with three setae mesally, posterior field of setae as illustrated ( Fig. 4 ).

Measurements. FW: 4800, HW: 3500, F: 1100, T: 1970, t1: 950, t2: 260, ctt1: 38, f1: 1040, f2: 920, f3: 720, Mx4: 370, IO: 300, D: 255, d: 292, IO/d: 1.45, PO: 0.85.

Specimens studied. Holotype male. COLOMBIA. Caldas, Manizales, Río Blanco Nature Reserve , 5º4’24.3”N: 75º26’16.2”W, 2592 m., 15.I.2011, A. N. García Aldrete and R. González . Paratypes: 1 male, same location and collectors, 5º3’56.8”N: 75º26’52.5”W, 2339 m., 16.I.2011, Beating dry vegetation . 1 male, Valle del Cauca, Sevilla, Kilómetro 14, 4º10’12.0”N: 75º51’28.7”W, 2170 m., 3.XI.2013. N. Hassy, on tree trunk . 1 male, Risaralda, Sanctuary of Flora and Fauna Otún Quimbaya, Cuchilla road, 4°43’N: 75°35’W, 2050 m., 20.III-30.IV.2004, Malaise trap .

Etymology. The specific epithet refers to the Colombian Andes mountain range, where, together with the holotype, the other specimens were collected.

Figures

FIGURES 1‒6. Goja andina n. sp. Male. 1. Forewing. 2. Hindwing. 3. Front view of head. 4. Left paraproct and epiproct. 5. Hypandrium. 6. Phallosome. Scales in mm.