Goja sierrajuarez, Carrejo & Obando & Casasola-González & García Aldrete, 2021

Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo & García Aldrete, Alfonso N., 2021, New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico, Zootaxa 5040 (4), pp. 451-481 : 468-469

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Goja sierrajuarez

n. sp.

Goja sierrajuarez   n. sp.

( Figs 25–33 View FIGURES 25‒33 )

Diagnosis. Male brachypterous, aedeagal arch slender, V-shaped, apex rectangular, hypandrium concave in the middle. Subgenital plate with sides converging to a round apex. Ninth sternum: spermapore circular, with elliptic, sclerotized sclerite, flanked by large, semi-elliptic pigmented areas.

Male. Color. Body pale yellowish. Head vertex dark ( Fig. 26 View FIGURES 25‒33 ), a pair of side dark brown spots partially surrounding the antennal sockets, with anterior ends ringed: a small, slightly pigmented spot on each gena. Compound eyes black. Ocelli absent, at most represented by dark brown small spots. Scape and pedicel pale yellow, flagellomeres posterior to f1 slightly more pigmented. Mx4 slightly more pigmented than the other palpomeres. Thorax dorsally brown, a pair of dark brown lateral stripes on each pleuron, extended to abdomen. Coxae and femora unpigmented. Tibiae and tarsomeres slightly more pigmented. Winglets ( Fig. 25 View FIGURES 25‒33 ) pigmented, with some brownish spots. Abdominal segments dorsally brown, terminal segments pigmented.

Morphology. Neotenic. Distal ends of labral sclerites not joined medially. Distal ends of f11 blunt. Lacinial tips narrow, outer cusp with five rounded denticles. Without coxal organ. Forewings as winglets, hindwings absent. Winglets ( Fig. 25 View FIGURES 25‒33 ) elongate, with several thick setae. Distal segment of hypandrium ( Fig. 28 View FIGURES 25‒33 ) with a narrow transverse band slightly pigmented; postero-lateral corners with fields of long setae on well developed lobes. Phallosome ( Fig. 27 View FIGURES 25‒33 ) closed anteriorly, side struts straight; aedeagal arch forming a nearly right angle, with a rectangular apical process short and sharp at sides; external parameres membranous; mesal sclerites broad, triangular, each with two small preapical prominences, connected to endophallus by sclerotic bands. Three pairs of short, acicular, central endophallic sclerites, supported by a membranous structure; first pair posterior, second pair anterior and third pair lateral. Epiproct ( Fig. 29 View FIGURES 25‒33 ) trapeziform, slightly pigmented, with field of setae posteriorly and one macrosetae mesally near anterior border. Paraprocts ( Fig. 29 View FIGURES 25‒33 ) with field of setae posteriorly; sensory fields with six to nine trichobothria, issuing from basal rosettes.

Measurements. FW: 471, F: 1292, T: 2029, t1: 942, t2: 228, ctt1: 34, Mx4: 383, f1: 1224, f2: 1040, f3: 663, f4: 500, f5: 287, f6: 296, f7: 232, f8: 247, f9: 182, f10: 202, f11: 224, IO: 337, D: 250, d: 209, IO/d: 1.61, PO: 0.84.

Female. Color. Head, thorax, legs, winglets ( Fig. 30 View FIGURES 25‒33 ) and abdomen same as the male.

Morphology. Compound eyes as in the male. Without ocelli. Coxal organ absent. Winglets elongate ( Fig. 30 View FIGURES 25‒33 ), with several thick setae. Subgenital plate ( Fig. 31 View FIGURES 25‒33 ) with slightly pigmented area, bilobed anteriorly; posterior border straight, with a few lateral long setae. Gonapophyses ( Fig. 33 View FIGURES 25‒33 ): v1 slender, acuminate, about 2/3 as long as v2+3; this robust, with a short proximal heel, distal process slightly bent, acuminate, armed with numerous microspines, v3 forming an elongate bulge on side of v2, with a row of eight or ten setae. Ninth sternum ( Fig. 33 View FIGURES 25‒33 ) almost circular, with a pair of large, tear-shaped areas on each side of longitudinal midline; spermapore small, distal, surrounded by a pigmented rim, with anterior end elongate and sharp. Posterior border of clunium bilobed over the area of the epiproct. Epiproct ( Fig. 32 View FIGURES 25‒33 ) slightly elongate, pigmented; anterior border convex, posterior border rounded, a field of macrosetae posteriorly, mesal macrosetae absent. Paraprocts ( Fig. 32 View FIGURES 25‒33 ) slightly elongate, pigmented, with field of macrosetae posteriorly, sensory fields with four to five trichobothria, issuing from basal rosettes.

Measurements. FW: 300, F: 1206, T: 1946, t1: 857, t2: 221, Mx4: 304, t1: 990, t2: 816, t3: 563, IO: 316, D: 247, d: 213, IO/d: 1.48, PO: 0.86.

Specimens studied. Holotype male. MEXICO. Oaxaca. Sierra Juárez, trail to El Relámpago , 9 km. S of La Esperanza, 5.III.2013, 17°35’31”N: 96°23’5.0”W, 2010 m. Sifting litter. A. N. García Aldrete and J. A. Casasola- González GoogleMaps   . Paratypes: 2 females, 4 nymphs ( CNIN), same data as the holotype GoogleMaps   .

Etymology. The specific epithet refers to the type locality, Sierra Juárez.


Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico