Ayacucho triarmatus, Benedetti & Pinto-da-Rocha, 2022

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

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Ayacucho triarmatus

nom. nov.

3.24. Ayacucho triarmatus nom. nov.

Figs 3E View Figure 3 , 7W, X View Figure 7 , 12K, L View Figure 12 , 28 View Figure 28

Tapacochana triseriata Roewer, 1959: 69 (desc.), fig. 1 (dorsal habitus and trochanter-femur IV); Kury 2003: 145 (cat.)


MALE: Measurements (n =1) DSW: 4.0; DSL: 4.4; CL: 1.4. FIVL: 4.2. ChL: 1.2. Coloration (in ethanol): Predominantly yellow with dark spots on the scutal areas I -IV, lateral and posterior margin of the dorsal scutum and free tergites. Dorsum: (Fig. 3E View Figure 3 ) Alpha-type DSS. Anterior margin with median elevation without granules distributed. Ocularium with a pair of spines, few granules. Carapace with granules distributed mainly in the lateral region of ocularium; posterior region with sparse granules. Areas I-IV with granules sparsely distributed; II-IV with some slightly larger granules. Posterior margin of dorsal scutum and free tergite I with a row of small blunt tubercles. Free tergites II-III with a row of acuminate tubercles, larger than those on free tergite I. Lateral margins of dorsal scutum with granules sparsely distributed. Chelicerae: (Fig. 3E View Figure 3 ) Not swollen. Segment I granulate. Segment II predominantly smooth, with 11 small teeth. Segment III with 12 teeth. Pedipalpus : Small granules distributed on the dorsal surface of the femur and patella. Trochanter with a large ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a row of four ventral setiferous tubercles, except at the apex. Tibia: prolateral IIi, retrolateral iIi. Tarsus: prolateral IIi, retrolateral IiIi. Venter: Coxae I-II with a middle row of six and eight small tubercles respectively; III-IV smooth. Three tiny tubercles among the apical part of coxae II-III and III-IV. Genital area smooth. Free sternites and anal operculum smooth. Legs: (Figs 3E View Figure 3 , 7W, X View Figure 7 ) Coxae I-II each one with an anterior and posterior apophysis; III with a prolateral apophysis; IV with setiferous granules distributed throughout its surface, and a proapical spiniform apophysis. Trochanters I-III smooth; IV with 3-4 dorsal median granules and a retroapical blunt tubercle. Femora I-III with granules scattered; IV granulate; a dorsal row of large acuminate tubercles, distributed in median region of the femur, with seven tubercles, the median tubercles larger than the basal ones; a retrolateral row of 12 large acuminate tubercles at along distal ⅓, with the size growing apically; a single apical retroventral acuminate tubercle; a proventral row of 11 acuminate tubercles along the distal ⅔, varying in size, interleaving several larger and smaller tubercles. Patellae and tibiae I-IV with sparse granulation, unarmed. Tarsal segmentation: (n =1) 6, 10,?, 6. Penis: (Fig. 12K, L View Figure 12 ) VP rectangular along the basal ½ and somewhat hexagonal along the apical ½ apical; wider at the middle region; distal margin concave; sinuous at lateral view. MS C1-C3 subapical short and straight; MS A1-A2 median short and straight (smaller than MS C). Lateral sacs long, apically slightly blunt, with long T3-like microsetae. Stylus with apex inflated and several small apical projections. Dorsal process conical and apically acuminated. Promontory slightly convex. - - FEMALE: unknown.


Similar to Ayacucho bambamarca comb. nov. and A. weyrauchi comb. nov. because three rows of spiniform tubercles in femur IV (Fig. 7W, X View Figure 7 ). Differs from A. weyrauchi comb. nov. by having nine spiniform tubercles in retrolateral row of femur IV (Fig. 7W, X View Figure 7 ); large tubercles in free tergites (Fig. 3E View Figure 3 ), distal margin of ventral plate slightly concave (Fig. 12K View Figure 12 ). Differs from A. bambamarca comb. nov. by having retrolateral row of tubercles covering only distal half distal of femur IV length (Fig. 7W, X View Figure 7 ); areas I-IV unarmed (Fig. 3E View Figure 3 ); distal margin of the VP without conspicuous lateral projections; dorsal process present in the penis (Fig. 12K, L View Figure 12 ).


Roewer described Cajamarca triseriata in 1957 and Tapacochana triseriata in 1959. Since they are considered here as belonging to the genus Ayacucho , Tapacochana triseriata Roewer, 1959 is a secondary homonym of Cajamarca triseriata Roewer, 1957 and must be replaced. C. triseriata Roewer, 1957 is considered here synonymy of Cajamarca bambamarca Roewer, 1957, but junior synonyms are also combinations with the genus where they are included and consequently compete for homonymy. Therefore, we create Ayacucho triarmatus nom. nov. as a replacement name for Roewer’s previous name.

Derivatio nominis.

The specific epithet, an adjective in nominative singular, formed by Latin prefix tri - + Latin armātus, ta, tum (armed), in reference to three rows of tubercles on male femur IV.


(Fig. 28 View Figure 28 ) PERU. Cajamarca . Cajamarca and near San Juan.

Material examined.

Type material: Holotype ♂, ' PERU, Cajamarca , near San Juan , between Chiclayo and Cajamarca , 1,900 m a.s.l., 06/VII/1956, Weyrauch leg. (SMF RII 12763 View Materials /34) . Additional material: 1 ♂, ' PERU, Cajamarca , Cajamarca , 3,200 m a.s.l., 6°40′S 78°45′W, 14/III/1958, Buschwald leg. (MUSM 0501238) GoogleMaps .














Ayacucho triarmatus

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo 2022

Tapacochana triseriata

Benedetti & Pinto-da-Rocha 2022