Cratogeomys Merriam 1895

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Geomyidae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press, pp. 859-870 : 859

Cratogeomys Merriam 1895

Cratogeomys Merriam 1895 , N. Amer. Fauna, 8: 150.

Type Species: Geomys merriami Thomas 1893

Synonyms: Craterogeomys J. A. Allen 1895 ; Platygeomys Merriam 1895 .

Species and subspecies: 8 species with 42 subspecies:

Species Cratogeomys castanops (Baird 1852)

Subspecies Cratogeomys castanops subsp. angusticeps Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. bullatus Russell and Baker 1955

Subspecies Cratogeomys castanops subsp. castanops Baird 1852

Subspecies Cratogeomys castanops subsp. clarkii (Baird 1855)

Subspecies Cratogeomys castanops subsp. consitus Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. dalquesti Hollander 1990

Subspecies Cratogeomys castanops subsp. excelsus Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. hirtus Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. jucundus Russell and Baker 1955

Subspecies Cratogeomys castanops subsp. parviceps Russell 1968

Subspecies Cratogeomys castanops subsp. perexiguus Russell 1968

Subspecies Cratogeomys castanops subsp. perplanus Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. pratensis Russell 1968

Subspecies Cratogeomys castanops subsp. sordidulus Russell and Baker 1955

Subspecies Cratogeomys castanops subsp. subsimus Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. surculus Russell 1968

Subspecies Cratogeomys castanops subsp. tamaulipensis Nelson and Goldman 1934

Subspecies Cratogeomys castanops subsp. ustulatus Russell and Baker 1955

Species Cratogeomys goldmani Merriam 1895

Subspecies Cratogeomys goldmani subsp. elibatus Russell 1968

Subspecies Cratogeomys goldmani subsp. goldmani Merriam 1895

Subspecies Cratogeomys goldmani subsp. maculats Alvarez and Alvarez-Castañeda 1996

Subspecies Cratogeomys goldmani subsp. peridoneus Nelson and Goldman 1934

Subspecies Cratogeomys goldmani subsp. planifrons Nelson and Goldman 1934

Subspecies Cratogeomys goldmani subsp. rubellus Nelson and Goldman 1934

Subspecies Cratogeomys goldmani subsp. subnubilus Nelson and Goldman 1934

Species Cratogeomys fumosus (Merriam 1892)

Species Cratogeomys gymnurus (Merriam 1892)

Subspecies Cratogeomys gymnurus subsp. gymnurus (Merriam 1892)

Subspecies Cratogeomys gymnurus subsp. imparilis (Goldman 1939)

Subspecies Cratogeomys gymnurus subsp. russelli Genoways and Jones 1969

Subspecies Cratogeomys gymnurus subsp. tellus Russell 1953

Species Cratogeomys merriami ( Thomas 1893)

Subspecies Cratogeomys merriami subsp. merriami ( Thomas 1894)

Subspecies Cratogeomys merriami subsp. estor (Merriam 1895)

Subspecies Cratogeomys merriami subsp. fulvescens (Merriam 1895)

Subspecies Cratogeomys merriami subsp. irolonis Nelson and Goldman 1934

Subspecies Cratogeomys merriami subsp. peraltus Goldman 1937

Subspecies Cratogeomys merriami subsp. perotensis (Merriam 1895)

Subspecies Cratogeomys merriami subsp. saccharalis Nelson and Goldman 1934

Species Cratogeomys neglectus (Merriam 1902)

Species Cratogeomys tylorhinus (Merriam 1895)

Subspecies Cratogeomys tylorhinus subsp. tylorhinus (Merriam 1895)

Subspecies Cratogeomys tylorhinus subsp. angustirostris (Merriam 1903)

Subspecies Cratogeomys tylorhinus subsp. atratus Russell 1953

Subspecies Cratogeomys tylorhinus subsp. brevirostris Russell 1968

Subspecies Cratogeomys tylorhinus subsp. planiceps (Merriam 1895)

Subspecies Cratogeomys tylorhinus subsp. zodius Russell 1953

Species Cratogeomys zinseri (Goldman 1939)

Discussion: Revised by Nelson and Goldman (1934 a) and Russell (1968 a, b), who included Cratogeomys as a valid subgenus of Pappogeomys , a position followed by Hall (1981) and Patton (1993). However, Honeycutt and Williams (1982), based on a phylogenetic analysis of allozyme data, and DeWalt et al. (1993 a) and Demastes et al. (2002), using mitochondrial DNA sequences, confirmed Russell’s hypothesis of the sister relationship between Pappogeomys and Cratogeomys , but treated the two as separate genera. While it remains a matter of personal opinion as to the distinction between genera and subgenera, I choose here to follow the current consensus of workers on this group (e.g., Lee and Baker, 1987; Davidow-Henry et al., 1989; Hollander, 1990). Russell (1968 b) recognized two species groups, the castanops group (with castanops and merriami ) and the gymnurus group (with fumosus , gymnurus , neglectus , tylorhinus , and zinseri ). While these groups have been confirmed by molecular phylogenetic studies (e.g., DeWalt et al., 1993 a; Demastes et al., 2002), both studies call into question the species boundaries recognized by Russell and earlier workers. Some changes are incorporated into the accounts below, but additional ones are likely following the completion of on-going studies by M. S. Hafner, J. W. Demastes, and co-workers (pers. Comm.). Hooper (1946) included Platygeomys within Cratogeomys . This name, with gymnurus as its type species, would be available for Russell’s gymnurus group, should that group warrant formal recognition. Key to eight species listed below given in León et al. (2001, Mammalian Species No. 685).