Ummidia funerea (Gertsch, 1936)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia funerea (Gertsch, 1936) Figs 21 , 22 , Map 3

Pachylomerus funereus Gertsch, 1936, 1; HOLOTYPE: 1 ♂ (IZC00327741) from Edinburg, Texas 26.3005 -98.1623 6, 25 m a.s.l., 1.vi.1935, coll. S Mulaik, deposited in AMNH, examined.

Pachylomerides celsus Gertsch & Mulaik, 1940, 313; HOLOTYPE: 1 ♂ (IZC327741) from 32miles SW of Laredo, Texas, United States, 5.viii.1935, coll. S Mulaik, deposited in AMNH, examined, syn. nov.

Ummidia celsa Brignoli, 1983, 117.

Diagnosis.

Ummidia funerea can be differentiated from all other species of Ummidia by having an anterior eye row which is extremely procurved, such that the posterior margin of the ALEs is even with the anterior margin of the AMEs and the ALEs are 2 × as large as the PLEs.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp, leg I and II Pt-Tr removed, in vial with specimen. General coloration. Carapace and chelicerae reddish black 10R 2.5/1, legs dark reddish brown 2.5YR 2.5/3. Abdomen very dark gray 10YR 3/1. Cephalothorax. Carapace 7.38 long, 7.17 wide. Pars cephalica 4.9 long. Foveal groove procurved, 0.5 long, 1.67 wide. Tubercle high. AER extremely procurved. PER slightly procurved. Eye group 1.14 long, 1.44 wide, AME 0.42, PME 0.22, ALE 0.62, PLE 0.34. Sternum with marginal fringe and few scattered setae, STRl 4.62, STRw 4.31. Chelicerae with anterior tooth row comprising six teeth, posterior margin with six teeth. Palpal endites with 22 cuspules spread over proximal half of endite face, lacking distal endite cuspules, ENDw 1.73, ENDl 3.05. Labium with nine cuspules, LBw 1.58, LBl 1.73. Rastellum with many spines. Abdomen setose, dorsally lighter with dorsodistal opalescence. Legs. F1 6.26; F1w 1.79; P1 3.37; Ti1 3.93; Mt1 2.93; Tr1 1.13; F3 4.61; F3w 2.12; P3 2.55; Ti3 2.48; Sd3 1.65; Mt3 2.99; Tr3 2.01; F4 6.01; F4w 2.02; P4 2.87; Ti4 3.78; Mt4 4.58; Tr4 2.03. Retrolateral face of tarsus IV with defined comb over proximal 2/3 of tarsus and paired spinules over distal third. Leg I spination pattern: TSp 0, TSpv 11, TSrd 1, TSr 0, TSrv 22, MtSp 6, MtSr 17, TrSp 7, TrSr 11. Pedipalps. PTl 3.38, PTw 1.37, Bl 3.26. Embolus relatively long, sinuous.

Variation, males

(n = 6). CL 6.08-9.1, 7.34 ± 0.45; CW 5.56-8.21, 6.84 ± 0.39; Cap 4.01-5.84, 4.82 ± 0.27; ENDl 0.83-1.19, 1.03 ± 0.07; ENDw 1.14-1.52, 1.37 ± 0.06; STRl 3.62-4.98, 4.31 ± 0.22; STRw 2.87-4.58, 3.82 ± 0.26; LBl 0.91-1.27, 1.08 ± 0.06; LBw 1.23-1.84, 1.5 ± 0.09; F1 5.22-7.31, 6.03 ± 0.32; F1w 1.54-2.02, 1.74 ± 0.08; P1 2.54-3.89, 3.12 ± 0.2; Ti1 3.2-4.79, 3.81 ± 0.22; Mt1 2.38-3.06, 2.7 ± 0.12; Tr1 0.87-1.34, 1.11 ± 0.06; F3 3.57-5.53, 4.47 ± 0.27; F3w 1.61-2.26, 1.95 ± 0.1; P3 2.02-3.08, 2.48 ± 0.15; Ti3 1.35-2.95, 2.27 ± 0.23; Mt3 2.35-3.27, 2.73 ± 0.15; Tr3 1.48-2.31, 1.86 ± 0.12; F4 4.78-7.14, 5.72 ± 0.35; F4w 1.53-2.18, 1.9 ± 0.1; P4 2.36-3.39, 2.79 ± 0.15; Ti4 3.07-4.26, 3.63 ± 0.16; Mt4 3.61-5.05, 4.24 ± 0.21; Tr4 1.57-2.24, 1.94 ± 0.09; TSp 0-8, 3.5 ± 1.31; TSpv 4-11, 6.67 ± 0.95; TSr 0-0, 0 ± 0; TSrv 19-28, 21.67 ± 1.36; PTl 3-4.19, 3.44 ± 0.16; PTw 1.09-1.52, 1.33 ± 0.06; BL 2.66-3.26, 3.06 ± 0.1.

Description of female exemplar.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. Left leg II and abdomen removed, in vial with specimen General coloration. Carapace and posterior legs dark yellowish brown 10YR 4/6, chelicerae and anterior legs dark reddish brown 2.5YR 2.5/4. Abdomen dark brown 10YR 3/3, spinnerets strong brown 7.5YR 4/6. Cephalothorax. Carapace 10.43 long, 9.83 wide. Pars cephalica 6.96 long. Foveal groove procurved, 0.78 long, 2.22 wide. Eye tubercle low, under ME. AER extremely procurved PER procurved. Eye group 1.42 long, 1.84 wide, AME 0.35, PME 0.42, ALE 0.82, PLE 0.34. Sternum sparsely setose around outer edges, thicker anteriorly, STRl 6.15, STRw 6.07. Chelicerae with anterior row comprising six teeth, posterior margin with six teeth. Palpal endites with 47 cuspules spread across proximal half of endite and 53 cuspules distally; equal in size to proximal cuspules anteriorly, smaller posteriorly, ENDw 2.2, ENDl 4.16. Labium with 15 cuspules, LBw 2.19, LBl 1.67. Rastellum with many strong spines on process and up cheliceral face. Abdomen. Evenly setose. Legs. F1 6.73; F1w 2.32; P1 4.15; Ti1 4.19; Mt1 2.72; Tr1 1.11; F3 5.26; F3w 2.88; P3 3.44; Ti3 3.2, Sd3 1.86; Mt3 2.93; Tr3 2.18; F4 6.55; F4w 2.72; P4 3.99; Ti4 3.95; Mt4 4.2; Tr4 1.97. Retrolateral face tarsus IV with defined tight comb with heavy setae ventrally. Pedipalps. PF 6.09, PP 3.36, PTi 3.9, PTr 3.27. Spermathecae curved medially, bulbs facing anteriorly.

Variation, females

(n = 7). CL 3.81-10.43, 7.35 ± 0.9; CW 3.28-9.83, 6.76 ± 0.88; Cap 2.5-6.96, 4.96 ± 0.62; ENDl 0.64-1.69, 1.14 ± 0.13; ENDw 0.76-1.84, 1.28 ± 0.13; STRl 2.32-6.18, 4.5 ± 0.56; STRw 2.18-6.07, 4.3 ± 0.55; LBl 0.55-1.67, 1.15 ± 0.16; LBw 0.87-2.19, 1.6 ± 0.18; F1 2.4-6.73, 4.77 ± 0.62; F1w 0.86-2.32, 1.66 ± 0.21; P1 1.54-4.15, 2.87 ± 0.37; Ti1 1.39-4.19, 2.95 ± 0.41; Mt1 1.06-2.98, 2.07 ± 0.27; Tr1 0.57-1.19, 0.92 ± 0.08; F3 1.19-5.4, 3.81 ± 0.59; F3w 1.1-2.91, 2.13 ± 0.26; P3 1.23-3.52, 2.52 ± 0.33; Ti3 1.09-3.2, 2.24 ± 0.31; Mt3 1.01-2.93, 2.06 ± 0.27; Tr3 0.79-2.18, 1.52 ± 0.19; F4 2.35-6.55, 4.79 ± 0.59; F4w 1.08-2.83, 2.06 ± 0.24; P4 1.46-3.99, 2.72 ± 0.35; Ti4 1.51-3.95, 2.85 ± 0.34; Mt4 1.35-4.24, 3.04 ± 0.42; Tr4 0.68-1.97, 1.37 ± 0.18; PF 2.13-6.09, 4.32 ± 0.57; PP 1.35-3.44, 2.48 ± 0.3; PTi 1.25-3.9, 2.73 ± 0.37; PTr 1.23-3.43, 2.39 ± 0.33.

Material examined.

United States: Texas: Brooks Co: 8 mi N Cantu , 31.9799 -99.9018 8, 540 m a.s.l. (UMM0313, 29.xii.1949, 1juv, AMNH) ; Cameron Co: Laguna Atascosa National Wildlife Refuge, 26.2238 -97.3345 1, 5 m a.s.l. (UMM0845, 15.v-4.vi.2010, 1♂, J King, E Riley, TAMU); Port Isabel , 26.0738 -97.207 5, 3 m a.s.l. (UMM0847, 8.iv.1992, 1♂, S Sweeten, TAMU) ; Hidalgo Co: Edinburg, 10 mi NW Edinburg, 26.3005 -98.1623 6, 25 m a.s.l. (IZC00327741, 1.vi.1935, 1♂, S Mulaik, AMNH); Texas, 34.9197 -99.9195 8, 540 m a.s.l. (UMM0262, 24.xii.1949, 1juv, Mulaik, AMNH); Edinburg , 26.3015 -98.1634 6, 30 m a.s.l. (UMM0267, 29.xii.1949, 1♀, S Mulaik, D Mulaik, AMNH); (UMM0321, ix.1936, 1♂, S Mulaik, AMNH); (UMM0614, ix.1936, 1♂, Mulaik, AMNH); (UMM0416, 1♂, Mulaik, AMNH) ; Starr Co: Just S of Hwy 83, 2.5 mi W of Sullivan City, 26.3015 -98.6334 5, 64 m a.s.l. (UMM0782, 6.ix.1974, 1juv, W Icenogle, AMNH); (UMM0783, 7.ix.1974, 1♀, AMNH); (UMM0784, 1juv, AMNH) ; Webb Co: Laredo, 27.5062 -99.5072 6, 128 m a.s.l. (UMM0422, 1♂, R Baird, AMNH); Near Hwy 83, 1.8 mi N junction with Hwy 35, 15 mi N Laredo, 27.7567 -99.4406 5, 212 m a.s.l. (UMM0430, 8.ix.1974, 1♀, W Icenogle, AMNH); (UMM0780, 9.ix.1974, 1♀, AMNH); (UMM0781, 1♀, AMNH) .

Figures

Map 3. Locality records for eastern United States species.

Figure 21. Ummidia funerea (Gertsch, 1936) from Edinburg, Texas. A male habitus illustration ICZ 327741 B female habitus illustration UMM 430. Scale bars: 4.0 mm.

Figure 22. Ummidia funerea (Gertsch, 1936) from Edinburg, Texas. A-D male holotype (ICZ 327741) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female exemplar (UMM 430). Scale bars: 1.0 mm.