Eutrichodesmus, Silvestri, 1910

Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species, ZooKeys 12 (12), pp. 1-46 : 41-44

publication ID

https://doi.org/ 10.3897/zookeys.12.167

publication LSID

lsid:zoobank.org:pub:B32EF8D5-C3C6-4C8C-8845-284125448425

DOI

https://doi.org/10.5281/zenodo.3792178

persistent identifier

https://treatment.plazi.org/id/936787D7-FF9B-FF9F-8ABB-4688FB92FD03

treatment provided by

Plazi

scientific name

Eutrichodesmus
status

 

Key to the species of Eutrichodesmus View in CoL

1 At least some metaterga with an evident mid-dorsal outgrowth or projection ( Figs 9 View Figure 9 A-D; 20A-D).................................................................................... 2

– All metaterga subequal, devoid of an evident mid-dorsal outgrowth or projection ( Figs 2 View Figure 2 A-F, 5A-D; 17A-D)................................................................... 9

2 Only last 3-8 metaterga in front of telson with a very evident mid-dorsal outgrowth. Body with 20 segments............................................................. 3

– Most of metaterga, including some of anterior body portion, with a high, often tuberculated projection ( Figs 9 View Figure 9 A-D; 20A-D). Body with 19 or 20 segments.......................................................................................................... 5

3 Metaterga 12-19 each with an increasingly evident, subtriangular, mid-dorsal outgrowth. Cave in Yunnan Prov., China.......................... E. dorsiangulatus View in CoL

– Only metaterga 16(17)-19 each with an evident, rather rounded, mid-dorsal outgrowth................................................................................................... 4

4 Paraterga narrower. Gonopod process dp less than one-quarter length of acropodite. North Vietnam.................................................. E. armatocaudatus View in CoL

– Paraterga broader. Gonopod process dp much longer, more than half length of acropodite. Cave in Yunnan Prov., China.......................... E. monodentus View in CoL

5 Mid-dorsal projections relatively low, not club-shaped, increasingly evident on metaterga 3–17(18) ( Fig. 20A View Figure 20 ) .............................................................. 6

– Mid-dorsal projections especially prominent, club-shaped, present on metaterga (4)5-19, only slightly lower on metatergum 19........................... 7

6 Body with 19 segments. Subtriangular mid-dorsal projections evident on metaterga 3-17, only slightly smaller on metatergum 18 ( Fig. 20A View Figure 20 ). Gonopod process dp rudimentary ( Figs 22E, F View Figure 22 ). Vietnam....... E. asteroides View in CoL sp. n.

– Body with 20 segments. Clearly tridentate mid-dorsal projections evident on metaterga 3-18, abruptly smaller on metatergum 19. Gonopod process dp small but evident. Malaysia.................................................... E. cavernicola View in CoL

7 Metatergum 4 with a high mid-dorsal projection ( Fig. 9A View Figure 9 ). Paraterga strongly declivous and quadrilobate ( Figs 9 View Figure 9 A-D, 10E). Gonopods as in Fig. 11 View Figure 11 . Vietnam........................................................................................ E. aster View in CoL sp. n.

– Mid-dorsal projections absent from metatergum 4. Paraterga less strongly declivous. Gonopods different. Malaysia......................................................... 8

8 Mid-dorsal projections on metaterga 5 and 6 straight in lateral view............. ................................................................................................. E. macclurei View in CoL

– Mid-dorsal projections on metaterga 5 and 6 slightly inclined anteriorly in lateral view................................................................................ E. reclinatus View in CoL

9 Body with 19 segments. Collum and metaterga virtually smooth, non-tuberculate, beset with long or short trichome ( Fig. 32 View Figure 32 A-F, 33C-E). Paraterga too short to allow complete body volvation ( Fig. 32A View Figure 32 ). Gonopod telopodite with a complex, lobiform dp ( Fig. 33F, G View Figure 33 ). Southwestern Pacific...................... 10

– Body with 20 segments. Collum and metaterga clearly tuberculate, trichome (if present) usually short ( Figs 25F View Figure 25 , 29F View Figure 29 ). Paraterga much broader, mostly reaching level of venter, so volvation usually complete. East and Southeast Asia........................................................................................................... 11

10 Collum and metaterga beset with long, 2-segmented, tactile setae. Tip of gonopod telopodite without teeth. Vanuatu.......................... E. communicans View in CoL

– Collum and metaterga beset with very short trichome ( Figs 32 View Figure 32 B-E, 33C-E). Tip of gonopod telopodite with a few teeth ( Figs 33F, G View Figure 33 ). Sulawesi, Indonesia..................................................................................... E. reductus View in CoL sp. n.

11 Paraterga mostly especially wide and only slightly declivous. Caves in Guangxi Province, China........................................................................................ 12

– Paraterga not so wide, strongly declivous ( Figs 2A View Figure 2 , 5A View Figure 5 , 13A, 17A, 28A) ... 13

12 Collum with a row of conspicuous teeth along fore edge of collum............... ...................................................................................................... E. similis View in CoL

– Collum with only minute teeth at fore edge of collum...................... E. latus View in CoL

13 Most metaterga with two transverse rows of bosses. Epiproct very strongly flattened dorsoventrally, subquadrate, spatuliform, with unincised margins. Gonopod telopodite particularly slender, about twice as long as coxa. Shikoku, Japan.......................................................................... E. peculiaris View in CoL

– Most metaterga with three transverse rows of bosses or conical tubercles. Epiproct never so strongly flattened. Gonopod telopodite usually shorter. Southeast Asia.................................................................................................... 14

14 Paraterga mostly set off laterally at base by a clear sulcus. Metaterga with three mixostictic rows of evident, conical tubercles, each surmounted by a long, 2-segmented seta. Gonopod telopodite with a particularly small, dentiform dp. Thailand.............................................................................. E. gremialis View in CoL

– Paraterga less markedly (if at all) set off laterally at base ( Figs 2A View Figure 2 , 5A View Figure 5 , 13A, 17A, 28A), largely continuing general outline of metaterga or nearly so (Figs 18D, 29C). Metatergal trichome very short or missing. Gonopod telopodite different.................................................................................................... 15

15 Paraterga relatively short, volvation apparently imperfect ( Fig. 24A View Figure 24 ). Body small, up to 6.5 mm long. Vietnam.......................................................... 16

– Paraterga broad enough to allow complete volvation (Figs 17A, 28A). Body usually larger............................................................................................. 17

16 Entire body pallid, up to 5.0 mm long. Gonopods simple ............. E. basalis View in CoL

– Vertex, collum, metaterga and much of telson grey, this strongly contrasting with pallid venter and legs ( Fig. 23 View Figure 23 ). Body 6.0- 6.5 mm long. Gonopods more elaborate ( Figs 26 View Figure 26 C-E)......................................................... E. griseus View in CoL sp. n.

17 Pattern of metatergal tuberculation isostictic ( Figs 5 View Figure 5 A-D). Seminal groove terminating on a very evident, pilose-spiculate pulvillus near base of a relatively shortened, lamellar acropodite ( Figs 6F, G View Figure 6 , 7 View Figure 7 A-F).... E. regularis View in CoL sp. n.

– Metatergal tuberculation mixostictic (Figs 13A-F, 24A). Seminal groove terminating distally to subapically, mostly without a pulvillus (seldom with a hairpad) (Figs 18G, H, 30) ....................................................................... 18

18 Distofemoral process of gonotelopodite bare, subunciform, pointed......... 19

– Distofemoral process of gonotelopodite nearly always conspicuously papillate (Figs 18G, H, 30), seldom redumentary (Figs 15A-F), never pointed ....... 20

19 Body ca 8.0 mm long. Gonopod tip bifid. Vietnam.................. E. demangei View in CoL

– Body ca 14.0 mm long. Gonopod tip simple, unciform. Cave in Yunnan Prov., China............................................................................ E. arcicollaris View in CoL

20 Paraterga mostly bilobate laterally and unilobate caudolaterally (Fig. 17A). Very small: 4.7-5.3 mm long and 0.65-0.7 mm wide. Gonopods as in Figs 18G, H. Vietnam......................................................... E. curticornis View in CoL sp. n.

– Paraterga mostly 3- or 4-lobate laterally (Figs 13A-D). Length ≥ 6.0 mm, width ≥ 1.1 mm. Gonopods different ....................................................... 21

21 Metaterga 18 and 19 not elevated dorsally. Caudolateral lobulations mostly deeply incised. Distofemoral process of gonopod bipartite. Caves in Guizhou Prov., China................................................................................... E. incisus View in CoL

– Metaterga 18 and 19 slightly elevated dorsally ( Figs 28 View Figure 28 A-F). Caudolateral lobulations not incised, tuberculiform. Distofemoral process of gonopod unipartite ( Fig. 30 View Figure 30 )......................................................................................... 22

22 Body 6.0- 6.5 mm long and 1.1-1.2 mm wide. Ozopores opening on top of porosteles representing penultimate of five lateral lobulations of respective paraterga, with caudolateral lobulations missing. Gonocoxae with only few macrosetae. Gonopods as in Fig. 30 View Figure 30 . Laos.................. E. multilobatus View in CoL sp. n.

– Body ≥ 8.0 mm long and 1.35 mm wide. Ozopores opening dorsally on caudalmost of 3-4 lateral lobulations of respective paraterga, caudolateral lobulations present. Gonocoxae with abundant macrosetae. Gonopods different.... ................................................................................................................. 23

23 Body 8.0- 8.5 mm long and 1.35-1.4 mm wide. Paraterga mostly 3-lobate ( Figs 2 View Figure 2 A-G). Gonopods as in Figs 3 View Figure 3 E-H. Cave in Guizhou Prov., China....... ....................................................................................... E. distinctus View in CoL sp. n.

– Body 12-13 mm long and 2.5-2.6 mm wide. Paraterga mostly 4-lobate (Figs 13A-D). Gonopods as in Figs 15A-F. Caves in Thanh Hoa Prov., Vietnam... ........................................................................................ E. filisetiger View in CoL sp. n.

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